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12D miR HR13_24

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Results for Pknox2_Pknox1

Z-value: 0.43

Motif logo

Transcription factors associated with Pknox2_Pknox1

Gene Symbol Gene ID Gene Info
ENSMUSG00000035934.9 Pbx/knotted 1 homeobox 2
ENSMUSG00000006705.6 Pbx/knotted 1 homeobox

Activity-expression correlation:

GenePromoterPearson corr. coef.P-valuePlot
Pknox1mm10_v2_chr17_+_31564749_315648540.567.5e-02Click!
Pknox2mm10_v2_chr9_-_37147257_371473230.402.2e-01Click!

Activity profile of Pknox2_Pknox1 motif

Sorted Z-values of Pknox2_Pknox1 motif

Promoter Log-likelihood Transcript Gene Gene Info
chr9_-_31913462 1.16 ENSMUST00000116615.3
BarH-like homeobox 2
chr1_-_87101590 0.50 ENSMUST00000113270.2
alkaline phosphatase, intestinal
chr19_-_9899450 0.46 ENSMUST00000025562.7
inner centromere protein
chr10_+_41810528 0.43 ENSMUST00000099931.3
sestrin 1
chr15_-_100599864 0.42 ENSMUST00000177247.2
ENSMUST00000177505.2
POU domain, class 6, transcription factor 1
chr1_+_87124946 0.40 ENSMUST00000044878.3
alkaline phosphatase 3, intestine, not Mn requiring
chr2_-_34913976 0.37 ENSMUST00000028232.3
PHD finger protein 19
chr2_+_162987502 0.36 ENSMUST00000117123.1
serum/glucocorticoid regulated kinase 2
chr2_+_162987330 0.34 ENSMUST00000018012.7
serum/glucocorticoid regulated kinase 2
chr17_-_70851710 0.33 ENSMUST00000166395.2
TGFB-induced factor homeobox 1
chr1_-_52817503 0.30 ENSMUST00000162576.1
inositol polyphosphate-1-phosphatase
chr2_-_31141802 0.30 ENSMUST00000073879.5
ENSMUST00000100208.2
ENSMUST00000100207.2
ENSMUST00000113555.1
ENSMUST00000075326.4
ENSMUST00000113552.2
ENSMUST00000136181.1
formin binding protein 1
chr11_+_78826575 0.30 ENSMUST00000147875.2
ENSMUST00000141321.1
LYR motif containing 9
chr7_-_131410325 0.30 ENSMUST00000154602.1
IKAROS family zinc finger 5
chr17_-_70851189 0.27 ENSMUST00000059775.8
TGFB-induced factor homeobox 1
chr2_-_31142002 0.25 ENSMUST00000113560.1
ENSMUST00000113564.2
ENSMUST00000113562.2
formin binding protein 1
chr5_-_100159261 0.25 ENSMUST00000139520.1
transmembrane protein 150C
chr8_+_20136455 0.24 ENSMUST00000179299.1
ENSMUST00000096485.4
predicted gene, 21811
chr7_-_48881032 0.23 ENSMUST00000058745.8
E2F transcription factor 8
chr5_-_134747241 0.22 ENSMUST00000015138.9
elastin
chr14_+_55575617 0.21 ENSMUST00000022826.5
fat storage-inducing transmembrane protein 1
chr8_+_56294552 0.19 ENSMUST00000034026.8
hydroxyprostaglandin dehydrogenase 15 (NAD)
chrX_+_49463926 0.19 ENSMUST00000130558.1
Rho GTPase activating protein 36
chr8_-_111522073 0.19 ENSMUST00000034437.6
ENSMUST00000038193.7
WD repeat domain 59
chr2_+_152911311 0.18 ENSMUST00000028970.7
myosin, light polypeptide kinase 2, skeletal muscle
chr9_+_44134562 0.18 ENSMUST00000034650.8
ENSMUST00000098852.2
melanoma cell adhesion molecule
chr10_-_123196916 0.17 ENSMUST00000020334.7
ubiquitin specific peptidase 15
chr10_-_41072279 0.16 ENSMUST00000061796.6
G protein-coupled receptor 6
chr4_+_11758147 0.16 ENSMUST00000029871.5
ENSMUST00000108303.1
cadherin 17
chr9_-_86880414 0.15 ENSMUST00000074501.5
ENSMUST00000098495.3
ENSMUST00000074468.6
ENSMUST00000036347.6
synaptosomal-associated protein 91
chr3_-_51560816 0.15 ENSMUST00000037141.7
SET domain containing (lysine methyltransferase) 7
chrX_+_13632769 0.15 ENSMUST00000096492.3
G protein-coupled receptor 34
chr2_-_113758638 0.14 ENSMUST00000099575.3
gremlin 1
chr7_-_45434590 0.14 ENSMUST00000107771.3
ENSMUST00000141761.1
RuvB-like protein 2
chr11_+_117986292 0.14 ENSMUST00000132676.1
phosphatidylglycerophosphate synthase 1
chr1_+_25830657 0.14 ENSMUST00000064487.1
predicted gene 9884
chr7_-_131410495 0.14 ENSMUST00000121033.1
ENSMUST00000046306.8
IKAROS family zinc finger 5
chr11_+_3202908 0.13 ENSMUST00000179770.1
ENSMUST00000110048.1
eukaryotic translation initiation factor 4E nuclear import factor 1
chr19_+_10018265 0.12 ENSMUST00000131407.1
RAB3A interacting protein (rabin3)-like 1
chr7_+_89404356 0.11 ENSMUST00000058755.3
frizzled homolog 4 (Drosophila)
chr14_-_30923547 0.11 ENSMUST00000170415.1
inter-alpha trypsin inhibitor, heavy chain 3
chr10_-_7792795 0.11 ENSMUST00000065124.1
RIKEN cDNA 6530403G13 gene
chr11_+_23256001 0.10 ENSMUST00000020538.6
ENSMUST00000109551.1
ENSMUST00000102870.1
ENSMUST00000102869.1
exportin 1, CRM1 homolog (yeast)
chr6_-_106800051 0.10 ENSMUST00000013882.7
ENSMUST00000049675.4
ENSMUST00000113239.3
cereblon
chr7_-_48881596 0.10 ENSMUST00000119223.1
E2F transcription factor 8
chr5_+_34761734 0.10 ENSMUST00000080036.2
huntingtin
chr5_+_150673739 0.10 ENSMUST00000016569.4
ENSMUST00000038900.8
PDS5, regulator of cohesion maintenance, homolog B (S. cerevisiae)
chr3_+_95232287 0.10 ENSMUST00000107200.1
ENSMUST00000107199.1
CDC42 small effector 1
chr6_+_54039558 0.10 ENSMUST00000046856.7
chimerin (chimaerin) 2
chrX_-_48034842 0.10 ENSMUST00000039026.7
apelin
chr15_+_59374198 0.09 ENSMUST00000079703.3
ENSMUST00000168722.1
non-SMC element 2 homolog (MMS21, S. cerevisiae)
chr5_+_138820080 0.09 ENSMUST00000179205.1
predicted gene 5294
chr1_+_74588347 0.09 ENSMUST00000113732.1
BCS1-like (yeast)
chr9_-_122294361 0.09 ENSMUST00000042546.2
anoctamin 10
chr3_+_116594959 0.09 ENSMUST00000029571.8
spindle assembly 6 homolog (C. elegans)
chr1_-_179546261 0.09 ENSMUST00000027769.5
transcription factor B2, mitochondrial
chr7_-_127273919 0.08 ENSMUST00000082428.3
selenophosphate synthetase 2
chr3_+_51415986 0.08 ENSMUST00000029303.7
N(alpha)-acetyltransferase 15, NatA auxiliary subunit
chr1_+_74236479 0.08 ENSMUST00000113820.2
ENSMUST00000006467.7
ENSMUST00000113819.1
actin related protein 2/3 complex, subunit 2
chr13_-_12106945 0.08 ENSMUST00000021750.7
ENSMUST00000170156.2
ryanodine receptor 2, cardiac
chr7_+_112225856 0.08 ENSMUST00000050149.5
ENSMUST00000106647.1
ENSMUST00000106648.1
microtubule associated monooxygenase, calponin and LIM domain containing 2
chr9_+_113930934 0.08 ENSMUST00000084885.5
ENSMUST00000009885.7
upstream binding protein 1
chr1_+_156558844 0.08 ENSMUST00000166172.2
ENSMUST00000027888.6
v-abl Abelson murine leukemia viral oncogene 2 (arg, Abelson-related gene)
chr1_+_74588289 0.08 ENSMUST00000113733.3
ENSMUST00000027358.4
BCS1-like (yeast)
chr11_-_80080928 0.08 ENSMUST00000103233.3
ENSMUST00000061283.8
cytokine receptor-like factor 3
chr13_+_109685994 0.08 ENSMUST00000074103.5
phosphodiesterase 4D, cAMP specific
chr15_-_76723814 0.07 ENSMUST00000036247.8
RIKEN cDNA C030006K11 gene
chr7_-_126584578 0.07 ENSMUST00000150311.1
ceroid lipofuscinosis, neuronal 3, juvenile (Batten, Spielmeyer-Vogt disease)
chr5_-_29478470 0.07 ENSMUST00000001608.7
ENSMUST00000165512.2
motor neuron and pancreas homeobox 1
chr5_+_34989473 0.07 ENSMUST00000114284.1
ENSMUST00000114285.1
regulator of G-protein signaling 12
chr16_-_96127604 0.07 ENSMUST00000133885.1
ENSMUST00000050884.7
high mobility group nucleosomal binding domain 1
chr6_+_54039935 0.07 ENSMUST00000114403.1
chimerin (chimaerin) 2
chr11_-_49712674 0.06 ENSMUST00000020624.6
ENSMUST00000145353.1
CCR4-NOT transcription complex, subunit 6
chr13_+_91461050 0.06 ENSMUST00000004094.8
ENSMUST00000042122.8
single-stranded DNA binding protein 2
chr3_+_85574109 0.06 ENSMUST00000127348.1
ENSMUST00000107672.1
ENSMUST00000107674.1
PET112 homolog (S. cerevisiae)
chr4_+_127021311 0.06 ENSMUST00000030623.7
splicing factor proline/glutamine rich (polypyrimidine tract binding protein associated)
chr2_+_146855861 0.06 ENSMUST00000099278.2
ENSMUST00000156232.1
polo-like kinase 1 substrate 1
chr11_-_98022594 0.06 ENSMUST00000103144.3
ENSMUST00000017552.6
ENSMUST00000092736.4
ENSMUST00000107562.1
calcium channel, voltage-dependent, beta 1 subunit
chr16_-_91069142 0.06 ENSMUST00000035689.1
ENSMUST00000114076.1
RIKEN cDNA 4932438H23 gene
chr18_-_79109391 0.05 ENSMUST00000025430.8
ENSMUST00000161465.2
SET binding protein 1
chr1_+_156558759 0.05 ENSMUST00000173929.1
v-abl Abelson murine leukemia viral oncogene 2 (arg, Abelson-related gene)
chr9_-_62980874 0.05 ENSMUST00000098651.4
protein inhibitor of activated STAT 1
chr19_+_10018193 0.05 ENSMUST00000113161.2
ENSMUST00000117641.1
RAB3A interacting protein (rabin3)-like 1
chr3_+_137864487 0.05 ENSMUST00000041045.7
H2A histone family, member Z
chr9_-_29963112 0.04 ENSMUST00000075069.4
neurotrimin
chr1_-_66945361 0.04 ENSMUST00000160100.1
myosin, light polypeptide 1
chr8_+_3655762 0.04 ENSMUST00000012849.8
ENSMUST00000169234.2
resistin
chr2_-_160872985 0.04 ENSMUST00000109460.1
ENSMUST00000127201.1
zinc fingers and homeoboxes 3
chr10_-_29535857 0.03 ENSMUST00000092623.3
R-spondin 3 homolog (Xenopus laevis)
chr8_-_69184177 0.03 ENSMUST00000185176.1
leucine zipper, putative tumor suppressor 1
chr17_-_48432723 0.03 ENSMUST00000046549.3
apolipoprotein B mRNA editing enzyme, catalytic polypeptide 2
chr8_+_121544378 0.03 ENSMUST00000181133.1
ENSMUST00000181679.1
RIKEN cDNA 1700030M09 gene
chr2_+_130012336 0.03 ENSMUST00000110299.2
transglutaminase 3, E polypeptide
chr10_+_50592669 0.03 ENSMUST00000035606.8
activating signal cointegrator 1 complex subunit 3
chr9_+_123366921 0.03 ENSMUST00000038863.7
leucyl-tRNA synthetase, mitochondrial
chr2_-_160872829 0.03 ENSMUST00000176141.1
zinc fingers and homeoboxes 3
chr4_+_141444654 0.03 ENSMUST00000006377.6
zinc finger and BTB domain containing 17
chr1_-_74588117 0.03 ENSMUST00000066986.6
zinc finger protein 142
chr16_-_11909398 0.03 ENSMUST00000127972.1
ENSMUST00000121750.1
ENSMUST00000096272.4
ENSMUST00000073371.6
calcineurin-like phosphoesterase domain containing 1
chr2_+_128967383 0.02 ENSMUST00000110320.2
ENSMUST00000110319.2
zinc finger CCCH type containing 6
chr8_+_19682268 0.02 ENSMUST00000153710.1
ENSMUST00000127799.1
predicted gene 6483
chr1_+_167618246 0.02 ENSMUST00000111380.1
retinoid X receptor gamma
chrX_-_109013389 0.02 ENSMUST00000033597.8
high-mobility group nucleosome binding domain 5
chr10_-_42276688 0.02 ENSMUST00000175881.1
ENSMUST00000056974.3
forkhead box O3
chr2_-_60673654 0.02 ENSMUST00000059888.8
ENSMUST00000154764.1
integrin beta 6
chr2_+_105682463 0.02 ENSMUST00000140173.1
ENSMUST00000135412.1
ENSMUST00000138365.1
ENSMUST00000145744.1
paired box gene 6
chr11_+_83964419 0.02 ENSMUST00000049714.8
ENSMUST00000092834.5
ENSMUST00000183714.1
ENSMUST00000183456.1
synergin, gamma
chr3_-_89402650 0.02 ENSMUST00000168325.1
ENSMUST00000057431.5
lens epithelial protein
chr5_+_93268247 0.01 ENSMUST00000121127.1
cyclin G2
chrX_-_102505359 0.01 ENSMUST00000087916.4
histone deacetylase 8
chr15_-_59374149 0.01 ENSMUST00000022976.4
RIKEN cDNA E430025E21 gene
chr8_-_115707778 0.01 ENSMUST00000109104.1
avian musculoaponeurotic fibrosarcoma (v-maf) AS42 oncogene homolog
chr11_-_46312220 0.01 ENSMUST00000129474.1
ENSMUST00000093166.4
ENSMUST00000165599.2
cytoplasmic FMR1 interacting protein 2
chr1_-_66945019 0.01 ENSMUST00000027151.5
myosin, light polypeptide 1
chr2_+_30416096 0.01 ENSMUST00000113601.3
ENSMUST00000113603.3
protein phosphatase 2A, regulatory subunit B (PR 53)
chr2_+_30416031 0.01 ENSMUST00000042055.3
protein phosphatase 2A, regulatory subunit B (PR 53)
chr2_-_76982455 0.01 ENSMUST00000011934.5
ENSMUST00000099981.2
ENSMUST00000099980.3
ENSMUST00000111882.2
ENSMUST00000140091.1
titin
chr10_-_42276744 0.01 ENSMUST00000105502.1
ENSMUST00000105501.1
forkhead box O3
chr13_+_63815240 0.01 ENSMUST00000021926.5
ENSMUST00000095724.3
ENSMUST00000143449.1
ENSMUST00000067821.5
excision repair cross-complementing rodent repair deficiency, complementation group 6 like 2
chr19_-_44069526 0.01 ENSMUST00000170801.1
ER lipid raft associated 1
chr1_+_58973521 0.01 ENSMUST00000114296.1
ENSMUST00000027185.4
STE20-related kinase adaptor beta
chr17_-_35121173 0.01 ENSMUST00000174024.1
casein kinase 2, beta polypeptide
chr5_+_137553517 0.00 ENSMUST00000136088.1
ENSMUST00000139395.1
ENSMUST00000136565.1
ENSMUST00000149292.1
ENSMUST00000125489.1
actin-like 6B
chr16_+_91391721 0.00 ENSMUST00000160764.1
predicted gene 21970
chr17_-_35121030 0.00 ENSMUST00000174306.1
casein kinase 2, beta polypeptide
chr2_-_91649785 0.00 ENSMUST00000111333.1
zinc finger protein 408
chr8_+_55940453 0.00 ENSMUST00000000275.7
glycine receptor, alpha 3 subunit
chr17_-_35121213 0.00 ENSMUST00000025246.6
casein kinase 2, beta polypeptide
chr16_+_55973881 0.00 ENSMUST00000050248.8
ribosomal protein L24
chr5_-_100500592 0.00 ENSMUST00000149714.1
ENSMUST00000046154.5
lin-54 homolog (C. elegans)

Network of associatons between targets according to the STRING database.

First level regulatory network of Pknox2_Pknox1

PNG image of the network

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Gene Ontology Analysis

Gene overrepresentation in biological process category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 1.2 GO:0042637 catagen(GO:0042637)
0.1 0.3 GO:0032877 positive regulation of DNA endoreduplication(GO:0032877)
0.1 0.1 GO:1900157 regulation of bone mineralization involved in bone maturation(GO:1900157)
0.1 0.4 GO:0061087 positive regulation of histone H3-K27 methylation(GO:0061087)
0.1 0.2 GO:0031446 regulation of fast-twitch skeletal muscle fiber contraction(GO:0031446) positive regulation of fast-twitch skeletal muscle fiber contraction(GO:0031448)
0.1 0.2 GO:0002314 germinal center B cell differentiation(GO:0002314)
0.0 0.1 GO:0090579 transcriptional activation by promoter-enhancer looping(GO:0071733) gene looping(GO:0090202) dsDNA loop formation(GO:0090579)
0.0 0.6 GO:0048387 negative regulation of retinoic acid receptor signaling pathway(GO:0048387)
0.0 0.1 GO:0061300 cerebellum vasculature development(GO:0061300)
0.0 0.1 GO:0035589 G-protein coupled purinergic nucleotide receptor signaling pathway(GO:0035589)
0.0 0.2 GO:0035616 histone H2B conserved C-terminal lysine deubiquitination(GO:0035616)
0.0 0.1 GO:1904580 regulation of intracellular mRNA localization(GO:1904580)
0.0 0.1 GO:0043396 corticotropin-releasing hormone secretion(GO:0043396) regulation of corticotropin-releasing hormone secretion(GO:0043397) positive regulation of corticotropin-releasing hormone secretion(GO:0051466) regulation of G-protein coupled receptor internalization(GO:1904020)
0.0 0.1 GO:0034184 positive regulation of maintenance of sister chromatid cohesion(GO:0034093) positive regulation of maintenance of mitotic sister chromatid cohesion(GO:0034184)
0.0 0.4 GO:1904262 negative regulation of TORC1 signaling(GO:1904262)
0.0 0.1 GO:2000812 response to rapamycin(GO:1901355) regulation of barbed-end actin filament capping(GO:2000812)
0.0 0.2 GO:0010890 positive regulation of sequestering of triglyceride(GO:0010890)
0.0 0.2 GO:0003376 sphingosine-1-phosphate signaling pathway(GO:0003376)
0.0 0.1 GO:0000720 pyrimidine dimer repair by nucleotide-excision repair(GO:0000720)
0.0 0.2 GO:0016185 synaptic vesicle budding from presynaptic endocytic zone membrane(GO:0016185)
0.0 0.1 GO:0001887 selenium compound metabolic process(GO:0001887) selenocysteine metabolic process(GO:0016259)
0.0 0.1 GO:0098735 positive regulation of the force of heart contraction(GO:0098735) positive regulation of calcium-transporting ATPase activity(GO:1901896)
0.0 0.2 GO:0018026 peptidyl-lysine monomethylation(GO:0018026)
0.0 0.1 GO:0032049 cardiolipin biosynthetic process(GO:0032049)
0.0 0.1 GO:0045876 positive regulation of sister chromatid cohesion(GO:0045876)
0.0 0.2 GO:0097033 respiratory chain complex III assembly(GO:0017062) mitochondrial respiratory chain complex III assembly(GO:0034551) mitochondrial respiratory chain complex III biogenesis(GO:0097033)
0.0 0.2 GO:0007567 parturition(GO:0007567)
0.0 0.1 GO:1900042 positive regulation of interleukin-2 secretion(GO:1900042)
0.0 0.1 GO:0038032 termination of G-protein coupled receptor signaling pathway(GO:0038032)
0.0 0.1 GO:0010735 positive regulation of transcription via serum response element binding(GO:0010735)
0.0 0.4 GO:0036342 post-anal tail morphogenesis(GO:0036342)
0.0 0.1 GO:0000056 ribosomal small subunit export from nucleus(GO:0000056)
0.0 0.1 GO:2000325 nuclear-transcribed mRNA catabolic process, no-go decay(GO:0070966) regulation of ligand-dependent nuclear receptor transcription coactivator activity(GO:2000325) positive regulation of ligand-dependent nuclear receptor transcription coactivator activity(GO:2000327)
0.0 0.0 GO:2000872 positive regulation of progesterone secretion(GO:2000872)
0.0 0.1 GO:0006390 transcription from mitochondrial promoter(GO:0006390)
0.0 0.1 GO:0017196 N-terminal peptidyl-methionine acetylation(GO:0017196)
0.0 0.1 GO:0035752 lysosomal lumen pH elevation(GO:0035752)

Gene overrepresentation in cellular component category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 0.2 GO:0071953 elastic fiber(GO:0071953)
0.1 0.6 GO:0061700 GATOR2 complex(GO:0061700)
0.1 0.5 GO:0000801 central element(GO:0000801)
0.1 0.2 GO:0098833 presynaptic endosome(GO:0098830) presynaptic endocytic zone(GO:0098833) presynaptic endocytic zone membrane(GO:0098835)
0.0 0.1 GO:0097255 R2TP complex(GO:0097255)
0.0 0.1 GO:0098536 deuterosome(GO:0098536)
0.0 0.1 GO:0030915 Smc5-Smc6 complex(GO:0030915)
0.0 0.4 GO:0035098 ESC/E(Z) complex(GO:0035098)
0.0 0.1 GO:0036195 muscle cell projection(GO:0036194) muscle cell projection membrane(GO:0036195)
0.0 0.1 GO:0031415 NatA complex(GO:0031415)

Gene overrepresentation in molecular function category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 0.9 GO:0004035 alkaline phosphatase activity(GO:0004035)
0.1 0.4 GO:0070728 leucine binding(GO:0070728)
0.1 0.3 GO:0004441 inositol-1,4-bisphosphate 1-phosphatase activity(GO:0004441)
0.1 0.2 GO:0035673 oligopeptide transmembrane transporter activity(GO:0035673)
0.0 0.2 GO:0004687 myosin light chain kinase activity(GO:0004687)
0.0 0.1 GO:0030297 transmembrane receptor protein tyrosine kinase activator activity(GO:0030297)
0.0 0.7 GO:0017081 chloride channel regulator activity(GO:0017081)
0.0 0.2 GO:0061649 ubiquitinated histone binding(GO:0061649)
0.0 0.1 GO:0000179 rRNA (adenine-N6,N6-)-dimethyltransferase activity(GO:0000179)
0.0 0.2 GO:0004957 prostaglandin E receptor activity(GO:0004957)
0.0 0.1 GO:0005219 ryanodine-sensitive calcium-release channel activity(GO:0005219)
0.0 0.1 GO:0043141 ATP-dependent 5'-3' DNA helicase activity(GO:0043141)
0.0 0.6 GO:0070410 co-SMAD binding(GO:0070410)
0.0 0.1 GO:0017169 CDP-alcohol phosphatidyltransferase activity(GO:0017169)
0.0 0.2 GO:0032050 clathrin heavy chain binding(GO:0032050)
0.0 0.2 GO:0038036 sphingosine-1-phosphate receptor activity(GO:0038036)
0.0 0.1 GO:0045028 G-protein coupled nucleotide receptor activity(GO:0001608) G-protein coupled purinergic nucleotide receptor activity(GO:0045028)
0.0 0.1 GO:0005049 nuclear export signal receptor activity(GO:0005049)
0.0 0.1 GO:0034452 dynactin binding(GO:0034452)

Gene overrepresentation in curated gene sets: REACTOME pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.0 0.6 REACTOME DOWNREGULATION OF SMAD2 3 SMAD4 TRANSCRIPTIONAL ACTIVITY Genes involved in Downregulation of SMAD2/3:SMAD4 transcriptional activity