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avrg: GFI1 WT vs 36n/n vs KD

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Results for Nkx2-5

Z-value: 1.22

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Transcription factors associated with Nkx2-5

Gene Symbol Gene ID Gene Info
Nkx2-5
ENSMUSG00000015579.6 NK2 homeobox 5

Activity-expression correlation:

GenePromoterPearson corr. coef.P-valuePlot
Nkx2-5mm39_v1_chr17_-_27060539_270605650.375.4e-01Click!

Activity profile of Nkx2-5 motif

Sorted Z-values of Nkx2-5 motif

Top targets:  

Promoter Log-likelihood Transcript Gene Gene Info
chr11_-_96868483 0.83 ENSMUST00000107624.8
Sp2
Sp2 transcription factor
chr14_+_51366512 0.82 ENSMUST00000095923.4
Rnase6
ribonuclease, RNase A family, 6
chr16_+_44764099 0.80 ENSMUST00000114622.10
ENSMUST00000166731.2
Cd200r3
CD200 receptor 3
chr15_-_82108531 0.78 ENSMUST00000109535.3
ENSMUST00000089161.10
Tnfrsf13c
tumor necrosis factor receptor superfamily, member 13c
chr4_+_85123654 0.72 ENSMUST00000030212.15
ENSMUST00000107189.8
ENSMUST00000107184.8
Sh3gl2
SH3-domain GRB2-like 2
chr7_+_127845984 0.70 ENSMUST00000164710.8
ENSMUST00000070656.12
Tgfb1i1
transforming growth factor beta 1 induced transcript 1
chr16_+_43993599 0.70 ENSMUST00000119746.8
ENSMUST00000088356.10
ENSMUST00000169582.3
Usf3
upstream transcription factor family member 3
chr3_-_129834788 0.70 ENSMUST00000168644.3
Sec24b
Sec24 related gene family, member B (S. cerevisiae)
chr17_+_69071546 0.62 ENSMUST00000233625.2
L3mbtl4
L3MBTL4 histone methyl-lysine binding protein
chr1_+_74752710 0.57 ENSMUST00000027356.7
Cyp27a1
cytochrome P450, family 27, subfamily a, polypeptide 1
chr14_+_51366306 0.57 ENSMUST00000226210.2
Rnase6
ribonuclease, RNase A family, 6
chr9_+_54824547 0.54 ENSMUST00000039742.8
Hykk
hydroxylysine kinase 1
chr11_+_32483290 0.53 ENSMUST00000102821.4
Stk10
serine/threonine kinase 10
chr8_+_125519751 0.53 ENSMUST00000041106.9
ENSMUST00000167588.9
Trim67
tripartite motif-containing 67
chr15_-_79212323 0.48 ENSMUST00000166977.9
Pla2g6
phospholipase A2, group VI
chr7_+_28869770 0.47 ENSMUST00000033886.8
ENSMUST00000209019.2
ENSMUST00000208330.2
Ggn
gametogenetin
chr9_+_44684248 0.41 ENSMUST00000128150.8
Ift46
intraflagellar transport 46
chr19_+_53131187 0.40 ENSMUST00000050096.15
ENSMUST00000237832.2
Add3
adducin 3 (gamma)
chr15_-_99355623 0.34 ENSMUST00000023747.14
Nckap5l
NCK-associated protein 5-like
chr15_-_5273645 0.34 ENSMUST00000120563.2
Ptger4
prostaglandin E receptor 4 (subtype EP4)
chr16_+_33504908 0.33 ENSMUST00000126532.2
Heg1
heart development protein with EGF-like domains 1
chr15_-_5273659 0.33 ENSMUST00000047379.15
Ptger4
prostaglandin E receptor 4 (subtype EP4)
chr14_+_21550921 0.33 ENSMUST00000182964.3
Kat6b
K(lysine) acetyltransferase 6B
chr19_-_59931432 0.32 ENSMUST00000170819.2
Rab11fip2
RAB11 family interacting protein 2 (class I)
chr9_+_44684285 0.31 ENSMUST00000239014.2
Ift46
intraflagellar transport 46
chr2_-_73722874 0.29 ENSMUST00000136958.8
ENSMUST00000112010.9
ENSMUST00000128531.8
ENSMUST00000112017.8
Atf2
activating transcription factor 2
chr7_+_127846121 0.25 ENSMUST00000167965.8
Tgfb1i1
transforming growth factor beta 1 induced transcript 1
chr9_-_53447908 0.25 ENSMUST00000150244.2
Atm
ataxia telangiectasia mutated
chr9_+_98305014 0.24 ENSMUST00000052068.11
Rbp1
retinol binding protein 1, cellular
chrX_+_164953444 0.24 ENSMUST00000130880.9
ENSMUST00000056410.11
ENSMUST00000096252.10
ENSMUST00000087169.11
Gemin8
gem nuclear organelle associated protein 8
chr4_+_62537750 0.24 ENSMUST00000084521.11
ENSMUST00000107424.2
Rgs3
regulator of G-protein signaling 3
chr5_+_16758777 0.24 ENSMUST00000030683.8
Hgf
hepatocyte growth factor
chr7_+_28869629 0.24 ENSMUST00000098609.4
Ggn
gametogenetin
chr4_-_93223746 0.23 ENSMUST00000066774.6
Tusc1
tumor suppressor candidate 1
chr2_+_173918089 0.23 ENSMUST00000155000.8
ENSMUST00000134876.8
ENSMUST00000147038.8
Stx16
syntaxin 16
chr6_+_92846338 0.22 ENSMUST00000113434.2
Gm15737
predicted gene 15737
chr3_-_107145968 0.22 ENSMUST00000197758.5
Prok1
prokineticin 1
chr11_-_101877832 0.21 ENSMUST00000107173.9
ENSMUST00000107172.8
Dusp3
dual specificity phosphatase 3 (vaccinia virus phosphatase VH1-related)
chr6_+_72281587 0.21 ENSMUST00000183018.8
ENSMUST00000182014.10
Sftpb
surfactant associated protein B
chr9_+_44684206 0.20 ENSMUST00000002099.11
Ift46
intraflagellar transport 46
chr2_-_73722932 0.20 ENSMUST00000154456.8
ENSMUST00000090802.11
ENSMUST00000055833.12
ENSMUST00000112007.8
ENSMUST00000112016.9
Atf2
activating transcription factor 2
chr6_+_47221293 0.20 ENSMUST00000199100.5
Cntnap2
contactin associated protein-like 2
chr5_+_16758897 0.19 ENSMUST00000196645.2
Hgf
hepatocyte growth factor
chrX_-_63320543 0.19 ENSMUST00000114679.2
ENSMUST00000069926.14
Slitrk4
SLIT and NTRK-like family, member 4
chr8_-_33374282 0.19 ENSMUST00000209107.4
ENSMUST00000209022.3
Nrg1
neuregulin 1
chr1_-_133589020 0.18 ENSMUST00000193504.6
ENSMUST00000195067.2
ENSMUST00000191896.6
ENSMUST00000194668.6
ENSMUST00000195424.6
ENSMUST00000179598.4
ENSMUST00000027736.13
Zc3h11a
Gm38394
Zc3h11a
zinc finger CCCH type containing 11A
predicted gene, 38394
zinc finger CCCH type containing 11A
chr17_+_27152276 0.18 ENSMUST00000237412.2
Phf1
PHD finger protein 1
chr1_+_172168764 0.18 ENSMUST00000056136.4
Kcnj10
potassium inwardly-rectifying channel, subfamily J, member 10
chr1_-_66856695 0.17 ENSMUST00000068168.10
ENSMUST00000113987.2
Kansl1l
KAT8 regulatory NSL complex subunit 1-like
chr5_+_124466146 0.17 ENSMUST00000111477.2
ENSMUST00000077376.3
2810006K23Rik
RIKEN cDNA 2810006K23 gene
chr12_+_91367764 0.16 ENSMUST00000021346.14
ENSMUST00000021343.8
Tshr
thyroid stimulating hormone receptor
chr6_-_115830307 0.16 ENSMUST00000032469.13
Mbd4
methyl-CpG binding domain protein 4
chr10_+_79824418 0.16 ENSMUST00000004784.11
ENSMUST00000105374.2
Cnn2
calponin 2
chr6_-_29380467 0.16 ENSMUST00000080428.13
Opn1sw
opsin 1 (cone pigments), short-wave-sensitive (color blindness, tritan)
chr5_-_124465875 0.15 ENSMUST00000184951.8
Mphosph9
M-phase phosphoprotein 9
chr2_-_32977182 0.14 ENSMUST00000102810.10
Garnl3
GTPase activating RANGAP domain-like 3
chr14_+_20724378 0.13 ENSMUST00000224492.2
ENSMUST00000223751.2
ENSMUST00000225108.2
ENSMUST00000224754.2
Sec24c
Sec24 related gene family, member C (S. cerevisiae)
chr7_-_126496534 0.13 ENSMUST00000120007.8
Tmem219
transmembrane protein 219
chr6_+_136530970 0.12 ENSMUST00000189535.2
Atf7ip
activating transcription factor 7 interacting protein
chr6_-_28421678 0.12 ENSMUST00000090511.4
Gcc1
golgi coiled coil 1
chr2_+_51928017 0.12 ENSMUST00000065927.6
Tnfaip6
tumor necrosis factor alpha induced protein 6
chr18_+_49965689 0.11 ENSMUST00000180611.8
Dmxl1
Dmx-like 1
chr2_-_73316809 0.11 ENSMUST00000141264.8
Wipf1
WAS/WASL interacting protein family, member 1
chr3_-_89067462 0.10 ENSMUST00000029686.4
Hcn3
hyperpolarization-activated, cyclic nucleotide-gated K+ 3
chr10_+_22520910 0.10 ENSMUST00000042261.5
Slc2a12
solute carrier family 2 (facilitated glucose transporter), member 12
chr6_-_136899167 0.10 ENSMUST00000032343.7
Erp27
endoplasmic reticulum protein 27
chr14_+_80237691 0.10 ENSMUST00000228749.2
ENSMUST00000088735.4
Olfm4
olfactomedin 4
chr6_+_57234937 0.09 ENSMUST00000228297.2
Vmn1r15
vomeronasal 1 receptor 15
chr6_+_127554931 0.09 ENSMUST00000071563.5
Cracr2a
calcium release activated channel regulator 2A
chr7_+_43339842 0.08 ENSMUST00000056329.7
Klk14
kallikrein related-peptidase 14
chr9_+_44684450 0.07 ENSMUST00000238800.2
ENSMUST00000147559.8
Ift46
intraflagellar transport 46
chr11_+_68858942 0.07 ENSMUST00000102606.10
ENSMUST00000018884.6
Slc25a35
solute carrier family 25, member 35
chr14_-_63509131 0.07 ENSMUST00000132122.2
Gata4
GATA binding protein 4
chr11_-_97909134 0.07 ENSMUST00000107561.9
Cacnb1
calcium channel, voltage-dependent, beta 1 subunit
chr15_-_79212400 0.07 ENSMUST00000173163.8
ENSMUST00000047816.15
ENSMUST00000172403.9
ENSMUST00000173632.8
Pla2g6
phospholipase A2, group VI
chr11_-_96807192 0.06 ENSMUST00000144731.8
ENSMUST00000127048.8
Cdk5rap3
CDK5 regulatory subunit associated protein 3
chr15_-_99355593 0.06 ENSMUST00000161948.2
Nckap5l
NCK-associated protein 5-like
chr6_-_115830249 0.06 ENSMUST00000122816.3
Mbd4
methyl-CpG binding domain protein 4
chr14_+_20724366 0.06 ENSMUST00000048657.10
Sec24c
Sec24 related gene family, member C (S. cerevisiae)
chr5_-_124465946 0.05 ENSMUST00000031344.13
Mphosph9
M-phase phosphoprotein 9
chr11_+_117223161 0.05 ENSMUST00000106349.2
Septin9
septin 9
chr12_-_115299134 0.05 ENSMUST00000195359.6
ENSMUST00000103530.3
Ighv1-59
immunoglobulin heavy variable V1-59
chr2_+_32498716 0.05 ENSMUST00000129165.2
St6galnac6
ST6 (alpha-N-acetyl-neuraminyl-2,3-beta-galactosyl-1,3)-N-acetylgalactosaminide alpha-2,6-sialyltransferase 6
chr19_+_16933471 0.04 ENSMUST00000087689.5
Prune2
prune homolog 2
chrX_-_164110372 0.03 ENSMUST00000058787.9
Glra2
glycine receptor, alpha 2 subunit
chr11_+_119205613 0.03 ENSMUST00000053245.7
Card14
caspase recruitment domain family, member 14
chr8_+_114369838 0.03 ENSMUST00000095173.3
ENSMUST00000034219.12
ENSMUST00000212269.2
Syce1l
synaptonemal complex central element protein 1 like
chr4_+_119494901 0.03 ENSMUST00000024015.3
Guca2a
guanylate cyclase activator 2a (guanylin)
chr14_-_78326435 0.03 ENSMUST00000118785.3
ENSMUST00000066437.5
Fam216b
family with sequence similarity 216, member B
chr5_-_8417982 0.03 ENSMUST00000088761.11
ENSMUST00000115386.8
ENSMUST00000050166.14
ENSMUST00000046838.14
ENSMUST00000115388.9
ENSMUST00000088744.12
ENSMUST00000115385.2
Adam22
a disintegrin and metallopeptidase domain 22
chr18_-_68562385 0.03 ENSMUST00000052347.8
Mc2r
melanocortin 2 receptor
chr7_-_97794679 0.02 ENSMUST00000098281.4
Omp
olfactory marker protein
chr12_-_115611981 0.02 ENSMUST00000103540.3
ENSMUST00000199266.2
Ighv8-12
immunoglobulin heavy variable V8-12
chrX_-_110372843 0.02 ENSMUST00000096348.10
ENSMUST00000113428.9
Rps6ka6
ribosomal protein S6 kinase polypeptide 6
chr6_+_30639217 0.02 ENSMUST00000031806.10
Cpa1
carboxypeptidase A1, pancreatic
chr3_+_97808506 0.01 ENSMUST00000029476.9
ENSMUST00000130620.2
ENSMUST00000122288.2
Sec22b
SEC22 homolog B, vesicle trafficking protein
chr8_-_84301457 0.01 ENSMUST00000238997.2
Tecr
trans-2,3-enoyl-CoA reductase
chr6_-_29380423 0.01 ENSMUST00000147483.3
Opn1sw
opsin 1 (cone pigments), short-wave-sensitive (color blindness, tritan)
chrX_-_110372800 0.01 ENSMUST00000137712.9
Rps6ka6
ribosomal protein S6 kinase polypeptide 6
chr14_+_53073315 0.01 ENSMUST00000197128.2
Trav7d-5
T cell receptor alpha variable 7D-5
chrX_-_110372733 0.01 ENSMUST00000065976.12
Rps6ka6
ribosomal protein S6 kinase polypeptide 6
chr9_+_44684324 0.01 ENSMUST00000214854.3
ENSMUST00000125877.8
Ift46
intraflagellar transport 46
chr6_+_30541581 0.01 ENSMUST00000096066.5
Cpa2
carboxypeptidase A2, pancreatic
chr11_+_87457479 0.01 ENSMUST00000239011.2
Septin4
septin 4
chrX_+_12937714 0.01 ENSMUST00000169594.9
ENSMUST00000089302.11
Usp9x
ubiquitin specific peptidase 9, X chromosome
chr1_+_74317709 0.01 ENSMUST00000077985.4
Gpbar1
G protein-coupled bile acid receptor 1
chr9_-_10904714 0.01 ENSMUST00000162484.8
ENSMUST00000160216.8
Cntn5
contactin 5
chr11_+_87457544 0.01 ENSMUST00000060360.7
Septin4
septin 4
chr3_+_3699205 0.01 ENSMUST00000108394.3
Hnf4g
hepatocyte nuclear factor 4, gamma
chrX_-_140508177 0.00 ENSMUST00000067841.8
Irs4
insulin receptor substrate 4
chr13_-_64645606 0.00 ENSMUST00000180282.2
Fam240b
family with sequence similarity 240 member B
chr3_-_57483330 0.00 ENSMUST00000120977.2
Wwtr1
WW domain containing transcription regulator 1
chr5_+_71815382 0.00 ENSMUST00000199967.5
Gabrb1
gamma-aminobutyric acid (GABA) A receptor, subunit beta 1
chr5_-_71815318 0.00 ENSMUST00000199357.2
Gabra4
gamma-aminobutyric acid (GABA) A receptor, subunit alpha 4
chr6_+_14901439 0.00 ENSMUST00000128567.8
Foxp2
forkhead box P2
chr12_-_114909863 0.00 ENSMUST00000103517.2
ENSMUST00000195417.2
Ighv1-43
immunoglobulin heavy variable V1-43
chr7_-_134100659 0.00 ENSMUST00000238294.2
D7Ertd443e
DNA segment, Chr 7, ERATO Doi 443, expressed

Network of associatons between targets according to the STRING database.

First level regulatory network of Nkx2-5

PNG image of the network

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Gene Ontology Analysis

Gene overrepresentation in biological process category:

Log-likelihood per target  Total log-likelihood Term Description
0.2 0.7 GO:2000388 negative regulation of integrin activation(GO:0033624) regulation of matrix metallopeptidase secretion(GO:1904464) matrix metallopeptidase secretion(GO:1990773) positive regulation of ovarian follicle development(GO:2000386) regulation of antral ovarian follicle growth(GO:2000387) positive regulation of antral ovarian follicle growth(GO:2000388) negative regulation of eosinophil migration(GO:2000417)
0.2 0.7 GO:1901301 regulation of cargo loading into COPII-coated vesicle(GO:1901301)
0.2 0.8 GO:0031296 B cell costimulation(GO:0031296)
0.1 0.5 GO:0032915 positive regulation of transforming growth factor beta2 production(GO:0032915)
0.1 1.0 GO:0030579 ubiquitin-dependent SMAD protein catabolic process(GO:0030579)
0.1 0.5 GO:0090154 positive regulation of sphingolipid biosynthetic process(GO:0090154) positive regulation of ceramide biosynthetic process(GO:2000304)
0.1 0.2 GO:0072434 signal transduction involved in G2 DNA damage checkpoint(GO:0072425) signal transduction involved in mitotic G2 DNA damage checkpoint(GO:0072434)
0.1 0.3 GO:0003017 lymph circulation(GO:0003017)
0.1 0.7 GO:1990416 cellular response to brain-derived neurotrophic factor stimulus(GO:1990416)
0.0 0.2 GO:0051935 amino acid neurotransmitter reuptake(GO:0051933) glutamate reuptake(GO:0051935)
0.0 1.4 GO:0019731 antibacterial humoral response(GO:0019731)
0.0 0.2 GO:0061086 negative regulation of histone H3-K27 methylation(GO:0061086)
0.0 0.2 GO:0006776 vitamin A metabolic process(GO:0006776)
0.0 1.0 GO:0001539 cilium or flagellum-dependent cell motility(GO:0001539) cilium-dependent cell motility(GO:0060285)
0.0 0.4 GO:0060665 regulation of branching involved in salivary gland morphogenesis by mesenchymal-epithelial signaling(GO:0060665)
0.0 0.2 GO:0045163 clustering of voltage-gated potassium channels(GO:0045163)
0.0 0.7 GO:0006491 N-glycan processing(GO:0006491)
0.0 0.1 GO:0060464 lung lobe formation(GO:0060464) diaphragm morphogenesis(GO:0060540)
0.0 0.2 GO:0072344 rescue of stalled ribosome(GO:0072344)
0.0 0.2 GO:1904587 response to glycoprotein(GO:1904587)
0.0 0.5 GO:2000060 positive regulation of protein ubiquitination involved in ubiquitin-dependent protein catabolic process(GO:2000060)
0.0 0.2 GO:0021842 directional guidance of interneurons involved in migration from the subpallium to the cortex(GO:0021840) chemorepulsion involved in interneuron migration from the subpallium to the cortex(GO:0021842)
0.0 0.2 GO:0090110 cargo loading into COPII-coated vesicle(GO:0090110)
0.0 0.2 GO:0018298 protein-chromophore linkage(GO:0018298)
0.0 0.1 GO:0032237 activation of store-operated calcium channel activity(GO:0032237)
0.0 0.3 GO:0007095 mitotic G2 DNA damage checkpoint(GO:0007095)

Gene overrepresentation in cellular component category:

Log-likelihood per target  Total log-likelihood Term Description
0.0 0.9 GO:0030127 COPII vesicle coat(GO:0030127)
0.0 0.1 GO:0043291 RAVE complex(GO:0043291)
0.0 0.1 GO:0044316 cone cell pedicle(GO:0044316)
0.0 0.7 GO:0030992 intraciliary transport particle B(GO:0030992)
0.0 0.2 GO:0097208 alveolar lamellar body(GO:0097208)
0.0 0.2 GO:0032797 SMN complex(GO:0032797)
0.0 0.7 GO:0035861 site of double-strand break(GO:0035861)
0.0 0.3 GO:0000786 nucleosome(GO:0000786)
0.0 0.2 GO:0000346 transcription export complex(GO:0000346)

Gene overrepresentation in molecular function category:

Log-likelihood per target  Total log-likelihood Term Description
0.2 1.0 GO:0048495 Roundabout binding(GO:0048495)
0.1 0.7 GO:0004957 prostaglandin E receptor activity(GO:0004957)
0.1 0.2 GO:0004677 DNA-dependent protein kinase activity(GO:0004677)
0.1 0.2 GO:0008263 pyrimidine-specific mismatch base pair DNA N-glycosylase activity(GO:0008263)
0.1 0.5 GO:0047499 calcium-independent phospholipase A2 activity(GO:0047499)
0.1 0.7 GO:0004571 mannosyl-oligosaccharide 1,2-alpha-mannosidase activity(GO:0004571)
0.0 0.5 GO:0008140 cAMP response element binding protein binding(GO:0008140)
0.0 0.2 GO:0004045 aminoacyl-tRNA hydrolase activity(GO:0004045)
0.0 0.2 GO:0015272 ATP-activated inward rectifier potassium channel activity(GO:0015272)
0.0 0.1 GO:0005222 intracellular cAMP activated cation channel activity(GO:0005222)
0.0 0.2 GO:0019841 retinol binding(GO:0019841)
0.0 0.2 GO:0008020 G-protein coupled photoreceptor activity(GO:0008020)
0.0 0.2 GO:0005176 ErbB-2 class receptor binding(GO:0005176)
0.0 0.2 GO:0033549 MAP kinase phosphatase activity(GO:0033549)
0.0 0.4 GO:0042056 chemoattractant activity(GO:0042056)
0.0 1.4 GO:0004540 ribonuclease activity(GO:0004540)

Gene overrepresentation in curated gene sets: canonical pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.0 1.2 PID RHOA PATHWAY RhoA signaling pathway
0.0 0.6 PID ARF6 PATHWAY Arf6 signaling events

Gene overrepresentation in curated gene sets: REACTOME pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 0.7 REACTOME PROSTANOID LIGAND RECEPTORS Genes involved in Prostanoid ligand receptors
0.0 0.9 REACTOME ANTIGEN PRESENTATION FOLDING ASSEMBLY AND PEPTIDE LOADING OF CLASS I MHC Genes involved in Antigen Presentation: Folding, assembly and peptide loading of class I MHC
0.0 0.6 REACTOME SYNTHESIS OF BILE ACIDS AND BILE SALTS VIA 24 HYDROXYCHOLESTEROL Genes involved in Synthesis of bile acids and bile salts via 24-hydroxycholesterol
0.0 0.7 REACTOME RETROGRADE NEUROTROPHIN SIGNALLING Genes involved in Retrograde neurotrophin signalling
0.0 0.5 REACTOME ACTIVATION OF THE AP1 FAMILY OF TRANSCRIPTION FACTORS Genes involved in Activation of the AP-1 family of transcription factors
0.0 0.5 REACTOME ACYL CHAIN REMODELLING OF PE Genes involved in Acyl chain remodelling of PE
0.0 0.2 REACTOME G2 M DNA DAMAGE CHECKPOINT Genes involved in G2/M DNA damage checkpoint
0.0 0.2 REACTOME BASE FREE SUGAR PHOSPHATE REMOVAL VIA THE SINGLE NUCLEOTIDE REPLACEMENT PATHWAY Genes involved in Base-free sugar-phosphate removal via the single-nucleotide replacement pathway
0.0 0.4 REACTOME IL 7 SIGNALING Genes involved in Interleukin-7 signaling
0.0 0.2 REACTOME ERKS ARE INACTIVATED Genes involved in ERKs are inactivated
0.0 0.2 REACTOME HORMONE LIGAND BINDING RECEPTORS Genes involved in Hormone ligand-binding receptors
0.0 0.2 REACTOME OPSINS Genes involved in Opsins