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GSE53960: rat RNA-Seq transcriptomic Bodymap

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Results for Atf5

Z-value: 0.56

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Transcription factors associated with Atf5

Gene Symbol Gene ID Gene Info
Atf5
ENSRNOG00000020060 activating transcription factor 5

Activity-expression correlation:

GenePromoterPearson corr. coef.P-valuePlot
Atf5rn6_v1_chr1_-_100810522_1008105220.026.7e-01Click!

Activity profile of Atf5 motif

Sorted Z-values of Atf5 motif

Top targets:  

Promoter Log-likelihood Transcript Gene Gene Info
chr16_-_39476384 21.37 ENSRNOT00000092968
Gpm6a
glycoprotein m6a
chr18_+_35574002 20.67 ENSRNOT00000093442
ENSRNOT00000070817
ENSRNOT00000093356
Myot
myotilin
chrX_-_29648359 20.36 ENSRNOT00000086721
ENSRNOT00000006777
Gpm6b
glycoprotein m6b
chr9_+_37727942 20.22 ENSRNOT00000016511
ENSRNOT00000074276
LOC100912306
myotilin-like
chr16_-_39476025 19.92 ENSRNOT00000014312
Gpm6a
glycoprotein m6a
chr10_-_8498422 16.93 ENSRNOT00000082332
Rbfox1
RNA binding protein, fox-1 homolog 1
chr10_+_90348299 13.37 ENSRNOT00000089616
Rundc3a
RUN domain containing 3A
chr1_+_106998623 12.40 ENSRNOT00000022383
Slc17a6
solute carrier family 17 member 6
chr9_-_60672246 10.93 ENSRNOT00000017761
Hecw2
HECT, C2 and WW domain containing E3 ubiquitin protein ligase 2
chr18_+_13386133 10.73 ENSRNOT00000020661
Asxl3
additional sex combs like 3, transcriptional regulator
chr6_+_27768943 10.72 ENSRNOT00000015820
Kif3c
kinesin family member 3C
chr7_-_118332577 10.63 ENSRNOT00000090370
Rbfox2
RNA binding protein, fox-1 homolog 2
chr1_+_192379543 10.09 ENSRNOT00000078705
Prkcb
protein kinase C, beta
chr9_-_28972835 9.58 ENSRNOT00000086967
ENSRNOT00000079684
Rims1
regulating synaptic membrane exocytosis 1
chr1_-_266074181 9.30 ENSRNOT00000026378
Psd
pleckstrin and Sec7 domain containing
chr10_+_86657285 8.37 ENSRNOT00000087346
Thra
thyroid hormone receptor alpha
chr20_-_5020150 7.95 ENSRNOT00000001146
Sapcd1
suppressor APC domain containing 1
chr16_-_7007287 7.28 ENSRNOT00000041216
Itih3
inter-alpha trypsin inhibitor, heavy chain 3
chr7_+_79638046 7.22 ENSRNOT00000029419
Zfpm2
zinc finger protein, multitype 2
chr1_+_245237736 6.91 ENSRNOT00000035814
Vldlr
very low density lipoprotein receptor
chr14_-_87701884 6.89 ENSRNOT00000079338
Mospd1
motile sperm domain containing 1
chr1_+_228684136 6.65 ENSRNOT00000028608
Olr337
olfactory receptor 337
chr2_-_220535751 6.51 ENSRNOT00000089082
Palmd
palmdelphin
chr12_+_41486076 6.47 ENSRNOT00000057242
Rita1
RBPJ interacting and tubulin associated 1
chr13_-_76049363 6.41 ENSRNOT00000075865
ENSRNOT00000007455
Brinp2
BMP/retinoic acid inducible neural specific 2
chr1_+_199495298 6.35 ENSRNOT00000086003
ENSRNOT00000026748
Itgad
integrin subunit alpha D
chr15_+_52557607 6.13 ENSRNOT00000019222
Gfra2
GDNF family receptor alpha 2
chr3_+_116899878 6.05 ENSRNOT00000090802
ENSRNOT00000066101
Sema6d
semaphorin 6D
chr9_-_71788281 6.03 ENSRNOT00000020078
Crygc
crystallin, gamma C
chr5_-_21345805 6.02 ENSRNOT00000081296
ENSRNOT00000007802
Car8
carbonic anhydrase 8
chr11_-_31180616 5.46 ENSRNOT00000065535
ENSRNOT00000052033
ENSRNOT00000002812
Synj1
synaptojanin 1
chr1_+_46873011 5.45 ENSRNOT00000068476
ENSRNOT00000067661
Synj2
synaptojanin 2
chr12_+_25036605 5.41 ENSRNOT00000001996
ENSRNOT00000084427
Limk1
LIM domain kinase 1
chr8_+_33239139 5.39 ENSRNOT00000011589
Arhgap32
Rho GTPase activating protein 32
chr13_+_43818010 5.09 ENSRNOT00000005532
LOC100909664
centrosomal protein of 170 kDa-like
chr8_+_52751854 4.93 ENSRNOT00000072518
Nxpe1
neurexophilin and PC-esterase domain family, member 1
chr1_-_198233215 4.77 ENSRNOT00000087928
Aldoa
aldolase, fructose-bisphosphate A
chr13_-_94859390 4.76 ENSRNOT00000005467
ENSRNOT00000079763
AABR07021840.1

chr1_-_57815038 4.66 ENSRNOT00000075401
Rgmb
repulsive guidance molecule family member B
chr1_+_198284473 4.37 ENSRNOT00000027022
Doc2a
double C2 domain alpha
chr9_-_45206427 4.06 ENSRNOT00000042227
Aff3
AF4/FMR2 family, member 3
chr2_+_216863428 3.83 ENSRNOT00000068413
Col11a1
collagen type XI alpha 1 chain
chr16_+_27399467 3.82 ENSRNOT00000065642
Tll1
tolloid-like 1
chr1_-_255815733 3.79 ENSRNOT00000047387
Cpeb3
cytoplasmic polyadenylation element binding protein 3
chr1_-_103380338 3.35 ENSRNOT00000019061
Mrgprx4
MAS related GPR family member X4
chr5_-_138462864 2.38 ENSRNOT00000011325
Ccdc30
coiled-coil domain containing 30
chr7_+_80750725 2.18 ENSRNOT00000079962
Oxr1
oxidation resistance 1
chrX_+_5549419 2.09 ENSRNOT00000004676
Fundc1
FUN14 domain containing 1
chr1_+_72399157 1.92 ENSRNOT00000022349
Zfp579
zinc finger protein 579
chr1_-_7480825 1.89 ENSRNOT00000048754
Phactr2
phosphatase and actin regulator 2
chr8_-_131899023 1.67 ENSRNOT00000034690
Zfp445
zinc finger protein 445
chr1_+_221872220 1.59 ENSRNOT00000028651
Nrxn2
neurexin 2
chr13_-_82758004 1.59 ENSRNOT00000003932
Atp1b1
ATPase Na+/K+ transporting subunit beta 1
chr1_-_169845487 1.46 ENSRNOT00000039989
Olr179
olfactory receptor 179
chr9_+_10044756 1.45 ENSRNOT00000071188
Pspn
persephin
chr3_+_75999481 1.44 ENSRNOT00000074642
Olr595
olfactory receptor 595
chr9_-_16643182 1.42 ENSRNOT00000024266
LOC108348250
cullin-7
chr1_+_229443102 1.39 ENSRNOT00000067320
Olr334
olfactory receptor 334
chr5_+_133864798 1.32 ENSRNOT00000091977
Tal1
TAL bHLH transcription factor 1, erythroid differentiation factor
chr3_-_79165698 1.31 ENSRNOT00000032617
Olr744
olfactory receptor 744
chr10_-_12674077 1.23 ENSRNOT00000041353
Olr1376
olfactory receptor 1376
chr10_-_12312692 1.22 ENSRNOT00000073059
Olr1355
olfactory receptor 1355
chr3_+_15688992 1.18 ENSRNOT00000045037
Olr395
olfactory receptor 395
chr19_+_9024451 1.09 ENSRNOT00000073744
LOC100910721
60S ribosomal protein L26-like
chr2_+_188141350 1.08 ENSRNOT00000078913
Gon4l
gon-4 like
chr13_-_76660043 1.06 ENSRNOT00000089332
Pappa2
pappalysin 2
chr1_+_198682230 1.02 ENSRNOT00000023995
Znf48
zinc finger protein 48
chr3_+_75768699 0.95 ENSRNOT00000071912
AABR07052798.1

chr6_-_99204825 0.93 ENSRNOT00000043602
Esr2
estrogen receptor 2
chr2_-_35192515 0.85 ENSRNOT00000086631
LOC100909937
olfactory receptor 149-like
chr1_-_89944752 0.84 ENSRNOT00000029680
Scgb2b24
secretoglobin, family 2B, member 24
chr1_+_68670127 0.79 ENSRNOT00000043828
Vom1r63
vomeronasal 1 receptor 63
chr6_-_99204590 0.65 ENSRNOT00000041872
ENSRNOT00000007109
Esr2
estrogen receptor 2
chr3_-_80601410 0.64 ENSRNOT00000022841
Atg13
autophagy related 13
chr7_-_121943121 0.55 ENSRNOT00000046460
Rpl26-ps1
ribosomal protein L26, pseudogene 1
chr3_+_15895191 0.54 ENSRNOT00000047879
Olr400
olfactory receptor 400
chr3_+_172510874 0.49 ENSRNOT00000073634
Nelfcd
negative elongation factor complex member C/D
chr1_-_31528083 0.39 ENSRNOT00000061041
Rpl26-ps2
ribosomal protein L26, pseudogene 2
chr13_-_76660245 0.25 ENSRNOT00000039210
Pappa2
pappalysin 2
chr4_+_60549197 0.24 ENSRNOT00000071249
ENSRNOT00000075621
Exoc4
exocyst complex component 4
chr17_+_45801528 0.16 ENSRNOT00000089221
Olr1664
olfactory receptor 1664
chr5_-_173209768 0.15 ENSRNOT00000045053
Atad3a
ATPase family, AAA domain containing 3A
chr3_-_73493621 0.14 ENSRNOT00000071782
Olr482
olfactory receptor 482
chr9_+_99735011 0.13 ENSRNOT00000051413
Olr1347
olfactory receptor 1347
chr1_+_200037749 0.10 ENSRNOT00000027630
Inpp5f
inositol polyphosphate-5-phosphatase F
chr3_+_15930518 0.06 ENSRNOT00000040288
Olr401
olfactory receptor 401
chr10_+_12602388 0.01 ENSRNOT00000060989
Olr1373
olfactory receptor 1373
chr3_+_16287056 0.01 ENSRNOT00000045825
Olr408
olfactory receptor 408
chr1_+_149963547 0.01 ENSRNOT00000042245
Olr20
olfactory receptor 20
chrM_+_2740 0.00 ENSRNOT00000047550
Mt-nd1
mitochondrially encoded NADH dehydrogenase 1

Network of associatons between targets according to the STRING database.

First level regulatory network of Atf5

PNG image of the network

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Gene Ontology Analysis

Gene overrepresentation in biological process category:

Log-likelihood per target  Total log-likelihood Term Description
4.1 20.4 GO:0051611 negative regulation of neurotransmitter uptake(GO:0051581) regulation of serotonin uptake(GO:0051611) negative regulation of serotonin uptake(GO:0051612)
3.4 10.1 GO:0035408 histone H3-T6 phosphorylation(GO:0035408)
3.2 9.6 GO:0031632 positive regulation of synaptic vesicle fusion to presynaptic membrane(GO:0031632)
2.8 8.4 GO:0017055 negative regulation of RNA polymerase II transcriptional preinitiation complex assembly(GO:0017055)
2.4 7.2 GO:0003221 right ventricular cardiac muscle tissue morphogenesis(GO:0003221)
1.7 6.9 GO:0034436 glycoprotein transport(GO:0034436)
1.6 12.4 GO:0098700 neurotransmitter loading into synaptic vesicle(GO:0098700)
1.6 9.3 GO:0006863 purine nucleobase transport(GO:0006863) nucleobase transport(GO:0015851)
1.5 10.6 GO:0010724 regulation of definitive erythrocyte differentiation(GO:0010724)
1.5 13.4 GO:0031284 positive regulation of guanylate cyclase activity(GO:0031284)
1.1 5.5 GO:0099590 neurotransmitter receptor internalization(GO:0099590)
1.0 6.1 GO:0035860 glial cell-derived neurotrophic factor receptor signaling pathway(GO:0035860)
1.0 41.3 GO:0051491 positive regulation of filopodium assembly(GO:0051491)
0.9 3.8 GO:0045900 negative regulation of translational elongation(GO:0045900)
0.7 6.4 GO:0010668 ectodermal cell differentiation(GO:0010668)
0.7 16.9 GO:2001014 regulation of skeletal muscle cell differentiation(GO:2001014)
0.5 1.6 GO:1903281 positive regulation of calcium:sodium antiporter activity(GO:1903281)
0.4 1.3 GO:0045799 basophil differentiation(GO:0030221) positive regulation of chromatin assembly or disassembly(GO:0045799) regulation of mast cell differentiation(GO:0060375)
0.4 3.8 GO:0035989 tendon development(GO:0035989)
0.4 1.6 GO:0008628 hormone-mediated apoptotic signaling pathway(GO:0008628)
0.4 5.4 GO:0046855 inositol phosphate dephosphorylation(GO:0046855)
0.3 4.8 GO:0030388 fructose 1,6-bisphosphate metabolic process(GO:0030388)
0.3 1.6 GO:0097116 gephyrin clustering involved in postsynaptic density assembly(GO:0097116)
0.2 10.9 GO:0030071 regulation of mitotic metaphase/anaphase transition(GO:0030071)
0.2 7.3 GO:0030212 hyaluronan metabolic process(GO:0030212)
0.2 2.2 GO:1902083 negative regulation of peptidyl-cysteine S-nitrosylation(GO:1902083)
0.2 5.4 GO:0051444 negative regulation of ubiquitin-protein transferase activity(GO:0051444)
0.2 6.5 GO:0045746 negative regulation of Notch signaling pathway(GO:0045746)
0.2 6.0 GO:0014912 negative regulation of smooth muscle cell migration(GO:0014912)
0.2 6.0 GO:0006730 one-carbon metabolic process(GO:0006730)
0.2 3.3 GO:0007635 chemosensory behavior(GO:0007635)
0.1 10.7 GO:0072384 organelle transport along microtubule(GO:0072384)
0.1 4.1 GO:0035116 embryonic hindlimb morphogenesis(GO:0035116)
0.1 4.4 GO:0048791 calcium ion-regulated exocytosis of neurotransmitter(GO:0048791)
0.1 6.0 GO:0002088 lens development in camera-type eye(GO:0002088)
0.1 4.2 GO:0071300 cellular response to retinoic acid(GO:0071300)
0.0 0.2 GO:0048341 paraxial mesoderm formation(GO:0048341)
0.0 4.7 GO:0030509 BMP signaling pathway(GO:0030509)
0.0 5.1 GO:0008360 regulation of cell shape(GO:0008360)
0.0 0.6 GO:0098780 response to mitochondrial depolarisation(GO:0098780)
0.0 2.1 GO:0061726 mitophagy(GO:0000422) mitochondrion disassembly(GO:0061726)
0.0 0.1 GO:0031161 phosphatidylinositol catabolic process(GO:0031161)
0.0 1.4 GO:0001658 branching involved in ureteric bud morphogenesis(GO:0001658)

Gene overrepresentation in cellular component category:

Log-likelihood per target  Total log-likelihood Term Description
1.3 10.7 GO:0071598 neuronal ribonucleoprotein granule(GO:0071598)
1.3 3.8 GO:0005592 collagen type XI trimer(GO:0005592)
1.2 9.6 GO:0098831 presynaptic active zone cytoplasmic component(GO:0098831)
1.1 5.4 GO:0031315 extrinsic component of mitochondrial outer membrane(GO:0031315)
1.0 46.7 GO:0044295 axonal growth cone(GO:0044295)
0.5 12.4 GO:0098563 integral component of synaptic vesicle membrane(GO:0030285) intrinsic component of synaptic vesicle membrane(GO:0098563)
0.4 6.9 GO:0034361 very-low-density lipoprotein particle(GO:0034361) triglyceride-rich lipoprotein particle(GO:0034385)
0.3 5.5 GO:0030132 clathrin coat of coated pit(GO:0030132)
0.2 1.4 GO:1990393 Cul7-RING ubiquitin ligase complex(GO:0031467) 3M complex(GO:1990393)
0.2 3.8 GO:1990124 messenger ribonucleoprotein complex(GO:1990124)
0.2 25.4 GO:0030018 Z disc(GO:0030018)
0.1 1.6 GO:0005890 sodium:potassium-exchanging ATPase complex(GO:0005890)
0.1 10.9 GO:0072686 mitotic spindle(GO:0072686)
0.1 0.5 GO:0032021 NELF complex(GO:0032021)
0.1 10.1 GO:0031526 brush border membrane(GO:0031526)
0.1 4.7 GO:0046658 anchored component of plasma membrane(GO:0046658)
0.1 0.6 GO:1990316 ATG1/ULK1 kinase complex(GO:1990316)
0.1 16.9 GO:0005802 trans-Golgi network(GO:0005802)
0.1 15.8 GO:0043197 dendritic spine(GO:0043197)
0.1 2.8 GO:0008305 integrin complex(GO:0008305)
0.1 2.1 GO:0031307 integral component of mitochondrial outer membrane(GO:0031307)
0.1 3.8 GO:0030672 synaptic vesicle membrane(GO:0030672) exocytic vesicle membrane(GO:0099501)
0.1 0.2 GO:0035748 myelin sheath abaxonal region(GO:0035748)
0.1 20.4 GO:0098857 membrane raft(GO:0045121) membrane microdomain(GO:0098857)
0.0 5.4 GO:0001650 fibrillar center(GO:0001650)
0.0 8.1 GO:0043025 neuronal cell body(GO:0043025)
0.0 5.5 GO:0048471 perinuclear region of cytoplasm(GO:0048471)

Gene overrepresentation in molecular function category:

Log-likelihood per target  Total log-likelihood Term Description
3.4 10.1 GO:0035403 histone kinase activity (H3-T6 specific)(GO:0035403)
2.3 6.9 GO:0034437 glycoprotein transporter activity(GO:0034437)
1.9 13.4 GO:0030250 cyclase activator activity(GO:0010853) guanylate cyclase activator activity(GO:0030250)
1.8 5.5 GO:0043812 phosphatidylinositol-4-phosphate phosphatase activity(GO:0043812)
0.8 5.4 GO:0004439 phosphatidylinositol-4,5-bisphosphate 5-phosphatase activity(GO:0004439)
0.8 12.4 GO:0005313 L-glutamate transmembrane transporter activity(GO:0005313)
0.8 9.3 GO:0015216 purine nucleotide transmembrane transporter activity(GO:0015216)
0.8 8.4 GO:0004887 thyroid hormone receptor activity(GO:0004887)
0.7 4.8 GO:0004332 fructose-bisphosphate aldolase activity(GO:0004332)
0.5 6.0 GO:0030215 semaphorin receptor binding(GO:0030215)
0.5 6.0 GO:0004089 carbonate dehydratase activity(GO:0004089)
0.5 20.7 GO:0051393 alpha-actinin binding(GO:0051393)
0.4 1.6 GO:1903924 estradiol binding(GO:1903924)
0.3 3.8 GO:0035613 RNA stem-loop binding(GO:0035613)
0.3 41.3 GO:0005262 calcium channel activity(GO:0005262)
0.3 6.0 GO:0005212 structural constituent of eye lens(GO:0005212)
0.2 4.7 GO:0015026 coreceptor activity(GO:0015026)
0.2 1.6 GO:0097109 neuroligin family protein binding(GO:0097109)
0.2 10.7 GO:0019894 kinesin binding(GO:0019894)
0.1 1.6 GO:0030955 potassium ion binding(GO:0030955)
0.1 27.6 GO:0003729 mRNA binding(GO:0003729) poly(A) RNA binding(GO:0044822)
0.1 11.8 GO:0031072 heat shock protein binding(GO:0031072)
0.1 9.6 GO:0017124 SH3 domain binding(GO:0017124)
0.1 1.9 GO:0004864 protein phosphatase inhibitor activity(GO:0004864)
0.1 4.4 GO:0005544 calcium-dependent phospholipid binding(GO:0005544)
0.1 7.3 GO:0004867 serine-type endopeptidase inhibitor activity(GO:0004867)
0.1 6.1 GO:0004896 cytokine receptor activity(GO:0004896)
0.1 10.2 GO:0016874 ligase activity(GO:0016874)
0.0 3.8 GO:0005518 collagen binding(GO:0005518)
0.0 0.1 GO:0034595 phosphatidylinositol phosphate 5-phosphatase activity(GO:0034595)
0.0 1.3 GO:0005201 extracellular matrix structural constituent(GO:0005201)
0.0 6.5 GO:0015631 tubulin binding(GO:0015631)
0.0 6.7 GO:0005549 odorant binding(GO:0005549)
0.0 1.3 GO:0000979 RNA polymerase II core promoter sequence-specific DNA binding(GO:0000979)
0.0 0.2 GO:0017160 Ral GTPase binding(GO:0017160)
0.0 3.4 GO:0035091 phosphatidylinositol binding(GO:0035091)

Gene overrepresentation in curated gene sets: canonical pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.7 10.1 PID TCR JNK PATHWAY JNK signaling in the CD4+ TCR pathway
0.2 6.4 PID INTEGRIN2 PATHWAY Beta2 integrin cell surface interactions
0.2 8.4 PID RXR VDR PATHWAY RXR and RAR heterodimerization with other nuclear receptor
0.2 6.9 PID LIS1 PATHWAY Lissencephaly gene (LIS1) in neuronal migration and development
0.1 5.5 PID EPHB FWD PATHWAY EPHB forward signaling
0.1 5.4 SIG REGULATION OF THE ACTIN CYTOSKELETON BY RHO GTPASES Genes related to regulation of the actin cytoskeleton
0.1 4.7 PID BMP PATHWAY BMP receptor signaling
0.1 5.4 PID TRKR PATHWAY Neurotrophic factor-mediated Trk receptor signaling
0.1 0.9 PID ERB GENOMIC PATHWAY Validated nuclear estrogen receptor beta network
0.1 4.8 PID HIF1 TFPATHWAY HIF-1-alpha transcription factor network
0.1 12.4 NABA ECM REGULATORS Genes encoding enzymes and their regulators involved in the remodeling of the extracellular matrix
0.0 2.0 PID SYNDECAN 1 PATHWAY Syndecan-1-mediated signaling events
0.0 7.9 NABA SECRETED FACTORS Genes encoding secreted soluble factors

Gene overrepresentation in curated gene sets: REACTOME pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.7 9.6 REACTOME NOREPINEPHRINE NEUROTRANSMITTER RELEASE CYCLE Genes involved in Norepinephrine Neurotransmitter Release Cycle
0.5 10.1 REACTOME TRAFFICKING OF GLUR2 CONTAINING AMPA RECEPTORS Genes involved in Trafficking of GluR2-containing AMPA receptors
0.3 10.7 REACTOME KINESINS Genes involved in Kinesins
0.3 6.0 REACTOME OTHER SEMAPHORIN INTERACTIONS Genes involved in Other semaphorin interactions
0.3 10.9 REACTOME SYNTHESIS OF PIPS AT THE PLASMA MEMBRANE Genes involved in Synthesis of PIPs at the plasma membrane
0.3 5.4 REACTOME SEMA3A PAK DEPENDENT AXON REPULSION Genes involved in Sema3A PAK dependent Axon repulsion
0.2 12.4 REACTOME AMINO ACID AND OLIGOPEPTIDE SLC TRANSPORTERS Genes involved in Amino acid and oligopeptide SLC transporters
0.2 6.9 REACTOME COLLAGEN FORMATION Genes involved in Collagen formation
0.1 9.9 REACTOME NUCLEAR RECEPTOR TRANSCRIPTION PATHWAY Genes involved in Nuclear Receptor transcription pathway
0.1 5.7 REACTOME NCAM1 INTERACTIONS Genes involved in NCAM1 interactions
0.1 4.8 REACTOME GLYCOLYSIS Genes involved in Glycolysis
0.1 4.7 REACTOME NETRIN1 SIGNALING Genes involved in Netrin-1 signaling
0.1 7.2 REACTOME FACTORS INVOLVED IN MEGAKARYOCYTE DEVELOPMENT AND PLATELET PRODUCTION Genes involved in Factors involved in megakaryocyte development and platelet production
0.1 1.6 REACTOME BASIGIN INTERACTIONS Genes involved in Basigin interactions
0.0 0.2 REACTOME INSULIN SYNTHESIS AND PROCESSING Genes involved in Insulin Synthesis and Processing