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Inflammatory response time course, HUVEC (Wada et al., 2009)

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Results for UUGUUCG

Z-value: 0.38

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miRNA associated with seed UUGUUCG

NamemiRBASE accession
hsa-miR-375
MIMAT0000728

Activity profile of UUGUUCG motif

Sorted Z-values of UUGUUCG motif

Top targets:  

Promoter Log-likelihood Transcript Gene Gene Info
chr12_+_20522179 1.13 ENST00000359062.3
PDE3A
phosphodiesterase 3A, cGMP-inhibited
chr13_-_72441315 1.01 ENST00000305425.4
ENST00000313174.7
ENST00000354591.4
DACH1
dachshund homolog 1 (Drosophila)
chr2_-_43453734 0.88 ENST00000282388.3
ZFP36L2
ZFP36 ring finger protein-like 2
chr10_-_131762105 0.79 ENST00000368648.3
ENST00000355311.5
EBF3
early B-cell factor 3
chr1_+_61547894 0.67 ENST00000403491.3
NFIA
nuclear factor I/A
chr6_-_16761678 0.65 ENST00000244769.4
ENST00000436367.1
ATXN1
ataxin 1
chr7_-_27183263 0.63 ENST00000222726.3
HOXA5
homeobox A5
chr18_+_42260861 0.61 ENST00000282030.5
SETBP1
SET binding protein 1
chr8_+_120220561 0.61 ENST00000276681.6
MAL2
mal, T-cell differentiation protein 2 (gene/pseudogene)
chr7_-_132261253 0.55 ENST00000321063.4
PLXNA4
plexin A4
chr1_-_100231349 0.53 ENST00000287474.5
ENST00000414213.1
FRRS1
ferric-chelate reductase 1
chr8_-_125384927 0.47 ENST00000297632.6
TMEM65
transmembrane protein 65
chr7_+_17338239 0.44 ENST00000242057.4
AHR
aryl hydrocarbon receptor
chr15_+_57210818 0.43 ENST00000438423.2
ENST00000267811.5
ENST00000452095.2
ENST00000559609.1
ENST00000333725.5
TCF12
transcription factor 12
chr22_+_51112800 0.42 ENST00000414786.2
SHANK3
SH3 and multiple ankyrin repeat domains 3
chr14_-_30396948 0.41 ENST00000331968.5
PRKD1
protein kinase D1
chr1_+_66999799 0.37 ENST00000371035.3
ENST00000371036.3
ENST00000371037.4
SGIP1
SH3-domain GRB2-like (endophilin) interacting protein 1
chr10_-_52383644 0.31 ENST00000361781.2
SGMS1
sphingomyelin synthase 1
chr17_+_38599693 0.31 ENST00000542955.1
ENST00000269593.4
IGFBP4
insulin-like growth factor binding protein 4
chr11_+_68080077 0.31 ENST00000294304.7
LRP5
low density lipoprotein receptor-related protein 5
chr3_+_16216137 0.29 ENST00000339732.5
GALNT15
UDP-N-acetyl-alpha-D-galactosamine:polypeptide N-acetylgalactosaminyltransferase 15
chr18_+_13218769 0.26 ENST00000399848.3
ENST00000361205.4
LDLRAD4
low density lipoprotein receptor class A domain containing 4
chr12_-_76425368 0.25 ENST00000602540.1
PHLDA1
pleckstrin homology-like domain, family A, member 1
chr1_+_2160134 0.25 ENST00000378536.4
SKI
v-ski avian sarcoma viral oncogene homolog
chr14_+_57735614 0.24 ENST00000261558.3
AP5M1
adaptor-related protein complex 5, mu 1 subunit
chr17_+_30813576 0.23 ENST00000313401.3
CDK5R1
cyclin-dependent kinase 5, regulatory subunit 1 (p35)
chr1_-_21377447 0.23 ENST00000374937.3
ENST00000264211.8
EIF4G3
eukaryotic translation initiation factor 4 gamma, 3
chr7_+_139026057 0.23 ENST00000541515.3
LUC7L2
LUC7-like 2 (S. cerevisiae)
chr10_-_126849068 0.21 ENST00000494626.2
ENST00000337195.5
CTBP2
C-terminal binding protein 2
chr20_+_306221 0.21 ENST00000342665.2
SOX12
SRY (sex determining region Y)-box 12
chrX_+_73641286 0.20 ENST00000587091.1
SLC16A2
solute carrier family 16, member 2 (thyroid hormone transporter)
chr11_+_12695944 0.20 ENST00000361905.4
TEAD1
TEA domain family member 1 (SV40 transcriptional enhancer factor)
chr2_+_204192942 0.20 ENST00000295851.5
ENST00000261017.5
ABI2
abl-interactor 2
chr11_+_33563821 0.19 ENST00000321505.4
ENST00000265654.5
ENST00000389726.3
KIAA1549L
KIAA1549-like
chr2_+_178257372 0.19 ENST00000264167.4
ENST00000409888.1
AGPS
alkylglycerone phosphate synthase
chr3_+_150126101 0.18 ENST00000361875.3
ENST00000361136.2
TSC22D2
TSC22 domain family, member 2
chr3_-_114790179 0.18 ENST00000462705.1
ZBTB20
zinc finger and BTB domain containing 20
chr18_-_53255766 0.18 ENST00000566286.1
ENST00000564999.1
ENST00000566279.1
ENST00000354452.3
ENST00000356073.4
TCF4
transcription factor 4
chr15_+_39542867 0.18 ENST00000318578.3
ENST00000561223.1
C15orf54
chromosome 15 open reading frame 54
chr15_-_101792137 0.17 ENST00000254190.3
CHSY1
chondroitin sulfate synthase 1
chr11_+_101981169 0.17 ENST00000526343.1
ENST00000282441.5
ENST00000537274.1
ENST00000345877.2
YAP1
Yes-associated protein 1
chr7_-_130080977 0.16 ENST00000223208.5
CEP41
centrosomal protein 41kDa
chr9_-_16870704 0.16 ENST00000380672.4
ENST00000380667.2
ENST00000380666.2
ENST00000486514.1
BNC2
basonuclin 2
chr19_-_14316980 0.15 ENST00000361434.3
ENST00000340736.6
LPHN1
latrophilin 1
chr8_-_92053212 0.15 ENST00000285419.3
TMEM55A
transmembrane protein 55A
chr1_+_180601139 0.14 ENST00000367590.4
ENST00000367589.3
XPR1
xenotropic and polytropic retrovirus receptor 1
chr9_-_135230336 0.14 ENST00000224140.5
ENST00000372169.2
ENST00000393220.1
SETX
senataxin
chr19_+_47421933 0.13 ENST00000404338.3
ARHGAP35
Rho GTPase activating protein 35
chr10_+_111967345 0.13 ENST00000332674.5
ENST00000453116.1
MXI1
MAX interactor 1, dimerization protein
chrX_-_18690210 0.13 ENST00000379984.3
RS1
retinoschisin 1
chr19_-_6279932 0.12 ENST00000252674.7
MLLT1
myeloid/lymphoid or mixed-lineage leukemia (trithorax homolog, Drosophila); translocated to, 1
chr7_+_157318477 0.12 ENST00000444154.1
AC006372.1
AC006372.1
chr13_-_31736027 0.12 ENST00000380406.5
ENST00000320027.5
ENST00000380405.4
HSPH1
heat shock 105kDa/110kDa protein 1
chr19_+_34919257 0.12 ENST00000246548.4
ENST00000590048.2
UBA2
ubiquitin-like modifier activating enzyme 2
chr13_+_73632897 0.11 ENST00000377687.4
KLF5
Kruppel-like factor 5 (intestinal)
chr14_-_70883708 0.11 ENST00000256366.4
SYNJ2BP
synaptojanin 2 binding protein
chr11_+_120207787 0.11 ENST00000397843.2
ENST00000356641.3
ARHGEF12
Rho guanine nucleotide exchange factor (GEF) 12
chr3_-_151176497 0.10 ENST00000282466.3
IGSF10
immunoglobulin superfamily, member 10
chr22_+_29168652 0.09 ENST00000249064.4
ENST00000444523.1
ENST00000448492.2
ENST00000421503.2
CCDC117
coiled-coil domain containing 117
chr1_-_213189108 0.09 ENST00000535388.1
ANGEL2
angel homolog 2 (Drosophila)
chr4_+_113152881 0.09 ENST00000274000.5
AP1AR
adaptor-related protein complex 1 associated regulatory protein
chr6_+_34433844 0.08 ENST00000244458.2
ENST00000374043.2
PACSIN1
protein kinase C and casein kinase substrate in neurons 1
chr3_+_152017181 0.08 ENST00000498502.1
ENST00000324196.5
ENST00000545754.1
ENST00000357472.3
MBNL1
muscleblind-like splicing regulator 1
chr9_+_26956371 0.07 ENST00000380062.5
ENST00000518614.1
IFT74
intraflagellar transport 74 homolog (Chlamydomonas)
chr1_-_205601064 0.07 ENST00000357992.4
ENST00000289703.4
ELK4
ELK4, ETS-domain protein (SRF accessory protein 1)
chr1_-_225840747 0.07 ENST00000366843.2
ENST00000366844.3
ENAH
enabled homolog (Drosophila)
chr5_+_138629417 0.07 ENST00000510056.1
ENST00000511249.1
ENST00000503811.1
ENST00000511378.1
MATR3
matrin 3
chr7_-_105517021 0.07 ENST00000318724.4
ENST00000419735.3
ATXN7L1
ataxin 7-like 1
chr22_+_40390930 0.06 ENST00000333407.6
FAM83F
family with sequence similarity 83, member F
chr19_-_18392422 0.06 ENST00000252818.3
JUND
jun D proto-oncogene
chr2_+_133174147 0.06 ENST00000329321.3
GPR39
G protein-coupled receptor 39
chr12_-_14133053 0.06 ENST00000609686.1
GRIN2B
glutamate receptor, ionotropic, N-methyl D-aspartate 2B
chr12_+_19592602 0.06 ENST00000398864.3
ENST00000266508.9
AEBP2
AE binding protein 2
chr14_-_55878538 0.05 ENST00000247178.5
ATG14
autophagy related 14
chr6_-_132834184 0.05 ENST00000367941.2
ENST00000367937.4
STX7
syntaxin 7
chr8_-_117768023 0.05 ENST00000518949.1
ENST00000522453.1
ENST00000521861.1
ENST00000518995.1
EIF3H
eukaryotic translation initiation factor 3, subunit H
chr1_+_199996702 0.04 ENST00000367362.3
NR5A2
nuclear receptor subfamily 5, group A, member 2
chr11_+_44117741 0.04 ENST00000395673.3
ENST00000343631.3
EXT2
exostosin glycosyltransferase 2
chr2_+_242641442 0.04 ENST00000313552.6
ENST00000406941.1
ING5
inhibitor of growth family, member 5
chr19_-_7293942 0.04 ENST00000341500.5
ENST00000302850.5
INSR
insulin receptor
chr13_+_31191920 0.04 ENST00000255304.4
USPL1
ubiquitin specific peptidase like 1
chr2_+_46926048 0.04 ENST00000306503.5
SOCS5
suppressor of cytokine signaling 5
chr19_+_16607122 0.04 ENST00000221671.3
ENST00000594035.1
ENST00000599550.1
ENST00000594813.1
C19orf44
chromosome 19 open reading frame 44
chr21_-_38362497 0.04 ENST00000427746.1
ENST00000336648.4
HLCS
holocarboxylase synthetase (biotin-(proprionyl-CoA-carboxylase (ATP-hydrolysing)) ligase)
chr7_+_75831181 0.03 ENST00000388802.4
ENST00000326382.8
SRRM3
serine/arginine repetitive matrix 3
chr1_+_62902308 0.03 ENST00000339950.4
USP1
ubiquitin specific peptidase 1
chr1_-_67896095 0.03 ENST00000370994.4
SERBP1
SERPINE1 mRNA binding protein 1
chr8_-_65711310 0.03 ENST00000310193.3
CYP7B1
cytochrome P450, family 7, subfamily B, polypeptide 1
chr7_+_139044621 0.03 ENST00000354926.4
C7orf55-LUC7L2
C7orf55-LUC7L2 readthrough
chr5_+_56111361 0.03 ENST00000399503.3
MAP3K1
mitogen-activated protein kinase kinase kinase 1, E3 ubiquitin protein ligase
chr16_+_69458428 0.03 ENST00000512062.1
ENST00000307892.8
CYB5B
cytochrome b5 type B (outer mitochondrial membrane)
chr4_+_155665123 0.03 ENST00000336356.3
LRAT
lecithin retinol acyltransferase (phosphatidylcholine--retinol O-acyltransferase)
chr2_-_172750733 0.03 ENST00000392592.4
ENST00000422440.2
SLC25A12
solute carrier family 25 (aspartate/glutamate carrier), member 12
chr14_-_96830207 0.02 ENST00000359933.4
ATG2B
autophagy related 2B
chr1_-_182573514 0.02 ENST00000367558.5
RGS16
regulator of G-protein signaling 16
chr8_-_38386175 0.02 ENST00000437935.2
ENST00000358138.1
C8orf86
chromosome 8 open reading frame 86
chr17_+_27369918 0.02 ENST00000323372.4
PIPOX
pipecolic acid oxidase
chr12_-_120806960 0.02 ENST00000257552.2
MSI1
musashi RNA-binding protein 1
chr10_+_114206956 0.01 ENST00000432306.1
ENST00000393077.2
VTI1A
vesicle transport through interaction with t-SNAREs 1A
chr5_+_145583156 0.01 ENST00000265271.5
RBM27
RNA binding motif protein 27
chr1_+_15853308 0.01 ENST00000375838.1
ENST00000375847.3
ENST00000375849.1
DNAJC16
DnaJ (Hsp40) homolog, subfamily C, member 16
chr9_+_4679555 0.01 ENST00000381858.1
ENST00000381854.3
CDC37L1
cell division cycle 37-like 1
chr3_+_137483579 0.01 ENST00000306087.1
SOX14
SRY (sex determining region Y)-box 14
chr10_+_102505468 0.01 ENST00000361791.3
ENST00000355243.3
ENST00000428433.1
ENST00000370296.2
PAX2
paired box 2
chr12_+_56401268 0.01 ENST00000262032.5
IKZF4
IKAROS family zinc finger 4 (Eos)
chr18_+_18943554 0.01 ENST00000580732.2
GREB1L
growth regulation by estrogen in breast cancer-like
chr12_+_103351444 0.01 ENST00000266744.3
ASCL1
achaete-scute family bHLH transcription factor 1
chr11_-_88796803 0.01 ENST00000418177.2
ENST00000455756.2
GRM5
glutamate receptor, metabotropic 5
chr5_+_174905398 0.01 ENST00000321442.5
SFXN1
sideroflexin 1
chr15_+_68346501 0.01 ENST00000249636.6
PIAS1
protein inhibitor of activated STAT, 1
chr9_+_15553055 0.01 ENST00000380701.3
CCDC171
coiled-coil domain containing 171
chr1_+_2985760 0.00 ENST00000378391.2
ENST00000514189.1
ENST00000270722.5
PRDM16
PR domain containing 16
chrX_+_24483338 0.00 ENST00000379162.4
ENST00000441463.2
PDK3
pyruvate dehydrogenase kinase, isozyme 3
chr10_+_45455207 0.00 ENST00000334940.6
ENST00000374417.2
ENST00000340258.5
ENST00000427758.1
RASSF4
Ras association (RalGDS/AF-6) domain family member 4
chr14_+_55493920 0.00 ENST00000395472.2
ENST00000555846.1
SOCS4
suppressor of cytokine signaling 4
chr17_+_55162453 0.00 ENST00000575322.1
ENST00000337714.3
ENST00000314126.3
AKAP1
A kinase (PRKA) anchor protein 1
chr7_+_92158083 0.00 ENST00000265732.5
ENST00000481551.1
ENST00000496410.1
RBM48
RNA binding motif protein 48
chr13_+_51483814 0.00 ENST00000336617.3
ENST00000422660.1
RNASEH2B
ribonuclease H2, subunit B

Network of associatons between targets according to the STRING database.

First level regulatory network of UUGUUCG

PNG image of the network

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Gene Ontology Analysis

Gene overrepresentation in biological process category:

Log-likelihood per target  Total log-likelihood Term Description
0.2 0.6 GO:0060435 bronchiole development(GO:0060435)
0.2 1.1 GO:0071321 cellular response to cGMP(GO:0071321)
0.2 0.5 GO:0021793 chemorepulsion of branchiomotor axon(GO:0021793)
0.1 1.0 GO:0060244 negative regulation of cell proliferation involved in contact inhibition(GO:0060244)
0.1 0.4 GO:0048170 positive regulation of long-term neuronal synaptic plasticity(GO:0048170)
0.1 0.9 GO:1904627 response to phorbol 13-acetate 12-myristate(GO:1904627) cellular response to phorbol 13-acetate 12-myristate(GO:1904628)
0.1 0.3 GO:0045210 FasL biosynthetic process(GO:0045210)
0.1 0.2 GO:0021722 superior olivary nucleus development(GO:0021718) superior olivary nucleus maturation(GO:0021722)
0.1 0.3 GO:0061304 retinal blood vessel morphogenesis(GO:0061304)
0.1 0.2 GO:0031064 negative regulation of histone deacetylation(GO:0031064)
0.1 0.4 GO:0089700 protein kinase D signaling(GO:0089700)
0.0 0.6 GO:0031580 membrane raft polarization(GO:0001766) membrane raft distribution(GO:0031580)
0.0 0.2 GO:0008611 ether lipid biosynthetic process(GO:0008611) glycerol ether biosynthetic process(GO:0046504) ether biosynthetic process(GO:1901503)
0.0 0.4 GO:2000253 positive regulation of feeding behavior(GO:2000253)
0.0 0.3 GO:0006686 sphingomyelin biosynthetic process(GO:0006686)
0.0 0.1 GO:0048203 vesicle targeting, trans-Golgi to endosome(GO:0048203)
0.0 0.0 GO:1902683 regulation of receptor localization to synapse(GO:1902683)
0.0 0.2 GO:0048386 positive regulation of retinoic acid receptor signaling pathway(GO:0048386)
0.0 0.2 GO:0060242 contact inhibition(GO:0060242)
0.0 0.3 GO:0010991 negative regulation of SMAD protein complex assembly(GO:0010991)
0.0 0.1 GO:0030950 establishment or maintenance of actin cytoskeleton polarity(GO:0030950)
0.0 0.2 GO:2000601 positive regulation of Arp2/3 complex-mediated actin nucleation(GO:2000601)
0.0 0.2 GO:0018095 protein polyglutamylation(GO:0018095)
0.0 0.7 GO:0060074 synapse maturation(GO:0060074)
0.0 0.2 GO:0070327 thyroid hormone transport(GO:0070327)
0.0 0.1 GO:1903750 regulation of intrinsic apoptotic signaling pathway in response to hydrogen peroxide(GO:1903750) negative regulation of intrinsic apoptotic signaling pathway in response to hydrogen peroxide(GO:1903751)
0.0 0.1 GO:0030643 cellular phosphate ion homeostasis(GO:0030643) cellular trivalent inorganic anion homeostasis(GO:0072502)
0.0 0.3 GO:0044342 positive regulation of insulin-like growth factor receptor signaling pathway(GO:0043568) type B pancreatic cell proliferation(GO:0044342)
0.0 0.1 GO:0032534 regulation of microvillus assembly(GO:0032534)
0.0 0.0 GO:0070781 response to biotin(GO:0070781)
0.0 0.1 GO:0048312 intracellular distribution of mitochondria(GO:0048312)
0.0 0.1 GO:0016240 autophagosome docking(GO:0016240)

Gene overrepresentation in cellular component category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 0.4 GO:0034751 aryl hydrocarbon receptor complex(GO:0034751)
0.1 0.3 GO:1990851 Wnt-Frizzled-LRP5/6 complex(GO:1990851)
0.1 0.6 GO:0042272 nuclear RNA export factor complex(GO:0042272)
0.1 0.4 GO:0071148 TEAD-1-YAP complex(GO:0071148)
0.1 0.2 GO:0016533 cyclin-dependent protein kinase 5 holoenzyme complex(GO:0016533)
0.0 0.3 GO:0000138 Golgi trans cisterna(GO:0000138)
0.0 0.5 GO:0002116 semaphorin receptor complex(GO:0002116)
0.0 0.1 GO:0031510 SUMO activating enzyme complex(GO:0031510)
0.0 0.2 GO:0097470 ribbon synapse(GO:0097470)
0.0 0.4 GO:0030122 AP-2 adaptor complex(GO:0030122)
0.0 0.1 GO:0097629 extrinsic component of omegasome membrane(GO:0097629)
0.0 0.2 GO:0031209 SCAR complex(GO:0031209)
0.0 0.2 GO:0016281 eukaryotic translation initiation factor 4F complex(GO:0016281)

Gene overrepresentation in molecular function category:

Log-likelihood per target  Total log-likelihood Term Description
0.4 1.1 GO:0004119 cGMP-inhibited cyclic-nucleotide phosphodiesterase activity(GO:0004119)
0.1 0.4 GO:0004874 aryl hydrocarbon receptor activity(GO:0004874)
0.1 0.6 GO:0034046 poly(G) binding(GO:0034046)
0.1 0.5 GO:0000293 ferric-chelate reductase activity(GO:0000293)
0.1 0.2 GO:0035033 histone deacetylase regulator activity(GO:0035033)
0.1 0.2 GO:0016534 cyclin-dependent protein kinase 5 activator activity(GO:0016534)
0.1 0.3 GO:0047493 sphingomyelin synthase activity(GO:0033188) ceramide cholinephosphotransferase activity(GO:0047493)
0.1 1.0 GO:0001075 transcription factor activity, RNA polymerase II core promoter sequence-specific binding involved in preinitiation complex assembly(GO:0001075)
0.1 0.4 GO:0030160 GKAP/Homer scaffold activity(GO:0030160)
0.0 0.1 GO:0001147 transcription termination site sequence-specific DNA binding(GO:0001147) transcription termination site DNA binding(GO:0001160)
0.0 0.2 GO:0050510 N-acetylgalactosaminyl-proteoglycan 3-beta-glucuronosyltransferase activity(GO:0050510)
0.0 0.8 GO:0035925 mRNA 3'-UTR AU-rich region binding(GO:0035925)
0.0 0.6 GO:0019911 structural constituent of myelin sheath(GO:0019911)
0.0 0.1 GO:0000822 inositol hexakisphosphate binding(GO:0000822)
0.0 0.3 GO:0071936 coreceptor activity involved in Wnt signaling pathway(GO:0071936)
0.0 0.3 GO:0031995 insulin-like growth factor II binding(GO:0031995)
0.0 0.2 GO:0030267 hydroxypyruvate reductase activity(GO:0016618) glyoxylate reductase (NADP) activity(GO:0030267)
0.0 0.5 GO:0017154 semaphorin receptor activity(GO:0017154)
0.0 0.1 GO:0035650 AP-1 adaptor complex binding(GO:0035650)
0.0 0.2 GO:0015349 thyroid hormone transmembrane transporter activity(GO:0015349)
0.0 0.1 GO:0001069 regulatory region RNA binding(GO:0001069)
0.0 0.1 GO:0034597 phosphatidylinositol-4,5-bisphosphate 4-phosphatase activity(GO:0034597)
0.0 0.4 GO:0035497 cAMP response element binding(GO:0035497)
0.0 0.4 GO:0004697 protein kinase C activity(GO:0004697)
0.0 0.1 GO:0044388 small protein activating enzyme binding(GO:0044388)
0.0 0.2 GO:0001087 transcription factor activity, sequence-specific DNA binding, RNA polymerase recruiting(GO:0001011) transcription factor activity, TFIIB-class binding(GO:0001087)
0.0 0.3 GO:0004653 polypeptide N-acetylgalactosaminyltransferase activity(GO:0004653)
0.0 0.4 GO:0070064 proline-rich region binding(GO:0070064)
0.0 0.0 GO:0050509 N-acetylglucosaminyl-proteoglycan 4-beta-glucuronosyltransferase activity(GO:0050509)

Gene overrepresentation in curated gene sets: canonical pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.0 0.6 PID WNT SIGNALING PATHWAY Wnt signaling network

Gene overrepresentation in curated gene sets: REACTOME pathways category:

Log-likelihood per target  Total log-likelihood Term Description
0.0 0.3 REACTOME REGULATION OF INSULIN LIKE GROWTH FACTOR IGF ACTIVITY BY INSULIN LIKE GROWTH FACTOR BINDING PROTEINS IGFBPS Genes involved in Regulation of Insulin-like Growth Factor (IGF) Activity by Insulin-like Growth Factor Binding Proteins (IGFBPs)
0.0 0.6 REACTOME MYOGENESIS Genes involved in Myogenesis