Motif ID: Barhl2

Z-value: 0.363


Transcription factors associated with Barhl2:

Gene SymbolEntrez IDGene Name
Barhl2 ENSMUSG00000034384.10 Barhl2

Activity-expression correlation:

Gene SymbolPromoterPearson corr. coef.P-valuePlot
Barhl2mm10_v2_chr5_-_106458440_106458470-0.156.2e-01Click!


Activity profile for motif Barhl2.

activity profile for motif Barhl2


Sorted Z-values histogram for motif Barhl2

Sorted Z-values for motif Barhl2



Network of associatons between targets according to the STRING database.



First level regulatory network of Barhl2

PNG image of the network

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Top targets:


Promoter Score Refseq Gene Symbol Gene Name
chrX_+_169879596 0.703 ENSMUST00000112105.1
ENSMUST00000078947.5
Mid1

midline 1

chr6_+_125552948 0.655 ENSMUST00000112254.1
ENSMUST00000112253.1
ENSMUST00000001995.7
Vwf


Von Willebrand factor homolog


chr12_-_34528844 0.605 ENSMUST00000110819.2
Hdac9
histone deacetylase 9
chr18_-_84086379 0.551 ENSMUST00000060303.8
Tshz1
teashirt zinc finger family member 1
chr19_+_8617991 0.485 ENSMUST00000010250.2
Slc22a6
solute carrier family 22 (organic anion transporter), member 6
chr3_-_80802789 0.484 ENSMUST00000107745.1
ENSMUST00000075316.4
Gria2

glutamate receptor, ionotropic, AMPA2 (alpha 2)

chr5_+_110839973 0.483 ENSMUST00000066160.1
Chek2
checkpoint kinase 2
chr18_-_78206408 0.472 ENSMUST00000163367.1
Slc14a2
solute carrier family 14 (urea transporter), member 2
chr11_+_85832551 0.432 ENSMUST00000000095.6
Tbx2
T-box 2
chr17_+_35439155 0.411 ENSMUST00000071951.6
ENSMUST00000078205.7
ENSMUST00000116598.3
ENSMUST00000076256.7
H2-Q7



histocompatibility 2, Q region locus 7



chr5_-_110839575 0.398 ENSMUST00000145318.1
Hscb
HscB iron-sulfur cluster co-chaperone homolog (E. coli)
chr18_+_61045139 0.379 ENSMUST00000025522.4
ENSMUST00000115274.1
Pdgfrb

platelet derived growth factor receptor, beta polypeptide

chr4_+_154960915 0.367 ENSMUST00000049621.6
Hes5
hairy and enhancer of split 5 (Drosophila)
chr17_-_49564262 0.362 ENSMUST00000057610.6
Daam2
dishevelled associated activator of morphogenesis 2
chr5_-_37824580 0.357 ENSMUST00000063116.9
Msx1
msh homeobox 1
chr1_-_163725123 0.345 ENSMUST00000159679.1
Mettl11b
methyltransferase like 11B
chr18_+_61639542 0.342 ENSMUST00000183083.1
ENSMUST00000183087.1
Gm20748

predicted gene, 20748

chr17_-_25727364 0.340 ENSMUST00000170070.1
ENSMUST00000048054.7
Chtf18

CTF18, chromosome transmission fidelity factor 18

chr18_-_80986578 0.337 ENSMUST00000057950.7
Sall3
sal-like 3 (Drosophila)
chr13_-_47106176 0.325 ENSMUST00000021807.6
ENSMUST00000135278.1
Dek

DEK oncogene (DNA binding)

chr2_-_127133909 0.320 ENSMUST00000110387.3
Ncaph
non-SMC condensin I complex, subunit H
chr1_+_165788681 0.303 ENSMUST00000161971.1
ENSMUST00000178336.1
ENSMUST00000005907.5
ENSMUST00000027849.4
Cd247



CD247 antigen



chr15_+_89568322 0.296 ENSMUST00000023295.2
Acr
acrosin prepropeptide
chr1_+_165788746 0.293 ENSMUST00000161559.2
Cd247
CD247 antigen
chr5_-_110839757 0.291 ENSMUST00000056937.5
Hscb
HscB iron-sulfur cluster co-chaperone homolog (E. coli)
chr6_-_101377897 0.280 ENSMUST00000075994.6
Pdzrn3
PDZ domain containing RING finger 3
chr2_+_181319714 0.274 ENSMUST00000098971.4
ENSMUST00000054622.8
ENSMUST00000108814.1
ENSMUST00000048608.9
ENSMUST00000108815.1
Rtel1




regulator of telomere elongation helicase 1




chr15_-_80014808 0.267 ENSMUST00000000500.6
Pdgfb
platelet derived growth factor, B polypeptide
chr9_-_13826946 0.264 ENSMUST00000147115.1
Cep57
centrosomal protein 57
chrX_+_71663665 0.260 ENSMUST00000070449.5
Gpr50
G-protein-coupled receptor 50
chr2_+_163225363 0.248 ENSMUST00000099110.3
ENSMUST00000165937.1
Tox2

TOX high mobility group box family member 2

chr9_-_13827029 0.245 ENSMUST00000148086.1
ENSMUST00000034398.5
Cep57

centrosomal protein 57

chr3_-_27153861 0.245 ENSMUST00000108300.1
ENSMUST00000108298.2
Ect2

ect2 oncogene

chr1_-_93343482 0.240 ENSMUST00000128253.1
Pask
PAS domain containing serine/threonine kinase
chr3_-_27153844 0.226 ENSMUST00000176242.2
ENSMUST00000176780.1
Ect2

ect2 oncogene

chr5_+_33658123 0.221 ENSMUST00000074849.6
ENSMUST00000079534.4
Tacc3

transforming, acidic coiled-coil containing protein 3

chr15_+_99074968 0.216 ENSMUST00000039665.6
Troap
trophinin associated protein
chr2_+_181319806 0.214 ENSMUST00000153112.1
Rtel1
regulator of telomere elongation helicase 1
chr8_+_57511833 0.213 ENSMUST00000067925.6
Hmgb2
high mobility group box 2
chr12_+_78748947 0.210 ENSMUST00000082024.5
Mpp5
membrane protein, palmitoylated 5 (MAGUK p55 subfamily member 5)
chr8_-_90908415 0.204 ENSMUST00000098517.1
Gm6658
predicted gene 6658
chr7_-_19715395 0.201 ENSMUST00000032555.9
ENSMUST00000093552.5
Tomm40

translocase of outer mitochondrial membrane 40 homolog (yeast)

chr5_+_135168382 0.199 ENSMUST00000111187.3
ENSMUST00000111188.1
Bcl7b

B cell CLL/lymphoma 7B

chr7_+_127841752 0.197 ENSMUST00000033075.7
Stx4a
syntaxin 4A (placental)
chr11_+_112782182 0.188 ENSMUST00000000579.2
Sox9
SRY-box containing gene 9
chr17_+_25727726 0.171 ENSMUST00000047273.1
Rpusd1
RNA pseudouridylate synthase domain containing 1
chr7_+_27486910 0.168 ENSMUST00000008528.7
Sertad1
SERTA domain containing 1
chr3_-_49757257 0.147 ENSMUST00000035931.7
Pcdh18
protocadherin 18
chr17_+_35220834 0.142 ENSMUST00000118793.1
Gm16181
predicted gene 16181
chr7_+_127841817 0.135 ENSMUST00000121705.1
Stx4a
syntaxin 4A (placental)
chr5_+_135168283 0.131 ENSMUST00000031692.5
Bcl7b
B cell CLL/lymphoma 7B
chr7_+_100495987 0.129 ENSMUST00000133044.1
Ucp2
uncoupling protein 2 (mitochondrial, proton carrier)
chr3_-_27153782 0.121 ENSMUST00000175857.1
ENSMUST00000177055.1
ENSMUST00000176535.1
Ect2


ect2 oncogene


chr2_+_156008088 0.115 ENSMUST00000088650.4
Ergic3
ERGIC and golgi 3
chr10_+_127290774 0.112 ENSMUST00000026475.8
ENSMUST00000139091.1
Ddit3

DNA-damage inducible transcript 3

chr16_-_22439719 0.110 ENSMUST00000079601.6
Etv5
ets variant gene 5
chr14_-_88471396 0.105 ENSMUST00000061628.5
Pcdh20
protocadherin 20
chr8_-_83594405 0.104 ENSMUST00000163837.1
Tecr
trans-2,3-enoyl-CoA reductase
chr2_+_156008045 0.104 ENSMUST00000006035.6
Ergic3
ERGIC and golgi 3
chr10_+_58394381 0.100 ENSMUST00000105468.1
Lims1
LIM and senescent cell antigen-like domains 1
chr2_+_118814237 0.094 ENSMUST00000028803.7
ENSMUST00000126045.1
Knstrn

kinetochore-localized astrin/SPAG5 binding

chr16_+_10835046 0.093 ENSMUST00000037913.8
Rmi2
RMI2, RecQ mediated genome instability 2, homolog (S. cerevisiae)
chr7_+_79660196 0.092 ENSMUST00000035977.7
Ticrr
TOPBP1-interacting checkpoint and replication regulator
chr1_-_153900198 0.090 ENSMUST00000123490.1
5830403L16Rik
RIKEN cDNA 5830403L16 gene
chr4_+_132768325 0.087 ENSMUST00000102561.4
Rpa2
replication protein A2
chrX_-_157415286 0.085 ENSMUST00000079945.4
ENSMUST00000138396.1
Phex

phosphate regulating gene with homologies to endopeptidases on the X chromosome (hypophosphatemia, vitamin D resistant rickets)

chr8_-_84104773 0.085 ENSMUST00000041367.7
Dcaf15
DDB1 and CUL4 associated factor 15
chr8_+_95081187 0.084 ENSMUST00000034239.7
Katnb1
katanin p80 (WD40-containing) subunit B 1
chr11_-_20332689 0.083 ENSMUST00000109594.1
Slc1a4
solute carrier family 1 (glutamate/neutral amino acid transporter), member 4
chr7_+_101969796 0.080 ENSMUST00000084852.5
Numa1
nuclear mitotic apparatus protein 1
chr8_-_107439585 0.078 ENSMUST00000077208.4
Rps26-ps1
ribosomal protein S26, pseudogene 1
chr11_-_120572822 0.077 ENSMUST00000168360.1
P4hb
prolyl 4-hydroxylase, beta polypeptide
chrX_-_59166080 0.075 ENSMUST00000119306.1
Fgf13
fibroblast growth factor 13
chr17_-_26939464 0.074 ENSMUST00000025027.8
ENSMUST00000114935.1
Cuta

cutA divalent cation tolerance homolog (E. coli)

chr7_-_3629874 0.073 ENSMUST00000155592.1
ENSMUST00000108641.3
Tfpt

TCF3 (E2A) fusion partner

chr14_+_55560480 0.071 ENSMUST00000121622.1
ENSMUST00000143431.1
ENSMUST00000150481.1
Dcaf11


DDB1 and CUL4 associated factor 11


chr5_-_115098295 0.071 ENSMUST00000100848.2
Gm10401
predicted gene 10401
chr7_+_25267669 0.070 ENSMUST00000169266.1
Cic
capicua homolog (Drosophila)
chr18_+_34736359 0.068 ENSMUST00000105038.2
Gm3550
predicted gene 3550
chr4_-_58553553 0.068 ENSMUST00000107575.2
ENSMUST00000107574.1
ENSMUST00000147354.1
Lpar1


lysophosphatidic acid receptor 1


chr18_+_42511496 0.067 ENSMUST00000025375.7
Tcerg1
transcription elongation regulator 1 (CA150)
chr15_-_100424092 0.067 ENSMUST00000154676.1
Slc11a2
solute carrier family 11 (proton-coupled divalent metal ion transporters), member 2
chr5_-_124249758 0.066 ENSMUST00000162812.1
Pitpnm2
phosphatidylinositol transfer protein, membrane-associated 2
chr11_-_20332654 0.063 ENSMUST00000004634.6
Slc1a4
solute carrier family 1 (glutamate/neutral amino acid transporter), member 4
chr17_+_27839974 0.062 ENSMUST00000071006.7
Snrpc
U1 small nuclear ribonucleoprotein C
chr8_-_83594440 0.059 ENSMUST00000019382.9
ENSMUST00000165740.1
Tecr

trans-2,3-enoyl-CoA reductase

chr2_-_58052832 0.058 ENSMUST00000090940.5
Ermn
ermin, ERM-like protein
chr10_+_80826656 0.058 ENSMUST00000060987.8
ENSMUST00000177850.1
ENSMUST00000180036.1
ENSMUST00000179172.1
Oaz1



ornithine decarboxylase antizyme 1



chr8_+_94214567 0.057 ENSMUST00000079961.6
Nup93
nucleoporin 93
chr14_-_31128924 0.057 ENSMUST00000064032.4
ENSMUST00000049732.5
ENSMUST00000090205.3
Smim4


small itegral membrane protein 4


chr19_-_6969746 0.054 ENSMUST00000025912.8
Plcb3
phospholipase C, beta 3
chr14_+_30825580 0.051 ENSMUST00000006701.5
Tmem110
transmembrane protein 110
chr7_+_82337218 0.043 ENSMUST00000173828.1
Adamtsl3
ADAMTS-like 3
chr11_-_120573253 0.043 ENSMUST00000026122.4
P4hb
prolyl 4-hydroxylase, beta polypeptide
chr19_-_29648355 0.042 ENSMUST00000159692.1
Ermp1
endoplasmic reticulum metallopeptidase 1
chr7_+_25301343 0.042 ENSMUST00000080288.3
Prr19
proline rich 19
chr5_-_134946917 0.037 ENSMUST00000051401.2
Cldn4
claudin 4
chr6_+_63255971 0.037 ENSMUST00000159561.1
ENSMUST00000095852.3
Grid2

glutamate receptor, ionotropic, delta 2

chr6_+_8259327 0.028 ENSMUST00000159378.1
Gm16039
predicted gene 16039
chr9_-_105131775 0.025 ENSMUST00000035179.6
Nudt16
nudix (nucleoside diphosphate linked moiety X)-type motif 16
chr14_+_21500879 0.023 ENSMUST00000182964.1
Kat6b
K(lysine) acetyltransferase 6B
chr1_-_171287931 0.022 ENSMUST00000111306.3
ENSMUST00000065941.5
Usp21

ubiquitin specific peptidase 21

chr7_+_113513829 0.020 ENSMUST00000033018.8
Far1
fatty acyl CoA reductase 1
chr6_-_95718800 0.020 ENSMUST00000079847.5
Suclg2
succinate-Coenzyme A ligase, GDP-forming, beta subunit
chr6_-_146502099 0.019 ENSMUST00000053273.8
Itpr2
inositol 1,4,5-triphosphate receptor 2
chr3_-_54714353 0.019 ENSMUST00000178832.1
Gm21958
predicted gene, 21958
chrX_+_82948861 0.017 ENSMUST00000114000.1
Dmd
dystrophin, muscular dystrophy
chr5_+_19907502 0.016 ENSMUST00000101558.3
Magi2
membrane associated guanylate kinase, WW and PDZ domain containing 2
chr11_+_116434087 0.013 ENSMUST00000057676.6
Ubald2
UBA-like domain containing 2
chr6_+_8259379 0.012 ENSMUST00000162034.1
ENSMUST00000160705.1
ENSMUST00000159433.1
Gm16039


predicted gene 16039


chr7_+_24370255 0.010 ENSMUST00000171904.1
Kcnn4
potassium intermediate/small conductance calcium-activated channel, subfamily N, member 4
chr18_+_4993795 0.010 ENSMUST00000153016.1
Svil
supervillin
chr1_+_105990652 0.008 ENSMUST00000118196.1
Zcchc2
zinc finger, CCHC domain containing 2
chr11_+_103966716 0.006 ENSMUST00000057921.3
ENSMUST00000063347.5
Arf2

ADP-ribosylation factor 2

chr1_-_171287916 0.006 ENSMUST00000111305.1
Usp21
ubiquitin specific peptidase 21
chr2_+_3513035 0.005 ENSMUST00000036350.2
Cdnf
cerebral dopamine neurotrophic factor
chr6_-_34317442 0.001 ENSMUST00000154655.1
ENSMUST00000102980.4
Akr1b3

aldo-keto reductase family 1, member B3 (aldose reductase)

chr6_+_125145235 0.000 ENSMUST00000119527.1
ENSMUST00000088276.6
ENSMUST00000051171.7
ENSMUST00000117675.1
Iffo1



intermediate filament family orphan 1




Gene Ontology Analysis

Gene overrepresentation in biological_process category:

Log-likelihood per target  Total log-likelihood Term Description
0.2 0.6 GO:0072223 metanephric glomerular mesangium development(GO:0072223) metanephric glomerular mesangial cell proliferation involved in metanephros development(GO:0072262)
0.2 0.6 GO:0060023 soft palate development(GO:0060023)
0.1 0.4 GO:0007521 muscle cell fate determination(GO:0007521) mammary placode formation(GO:0060596)
0.1 0.5 GO:1902990 mitotic telomere maintenance via semi-conservative replication(GO:1902990) negative regulation of t-circle formation(GO:1904430)
0.1 0.5 GO:1903463 mitotic cell cycle phase(GO:0098763) regulation of mitotic cell cycle DNA replication(GO:1903463)
0.1 0.4 GO:0035880 embryonic nail plate morphogenesis(GO:0035880)
0.1 0.3 GO:0002447 eosinophil activation involved in immune response(GO:0002278) eosinophil mediated immunity(GO:0002447) eosinophil degranulation(GO:0043308) regulation of eosinophil activation(GO:1902566)
0.1 0.3 GO:0007341 penetration of zona pellucida(GO:0007341)
0.1 0.4 GO:2000974 negative regulation of pro-B cell differentiation(GO:2000974)
0.1 0.3 GO:2001032 regulation of double-strand break repair via nonhomologous end joining(GO:2001032)
0.1 0.5 GO:0015840 urea transport(GO:0015840) urea transmembrane transport(GO:0071918)
0.1 0.2 GO:0035622 intrahepatic bile duct development(GO:0035622) renal vesicle induction(GO:0072034)
0.1 0.6 GO:0034983 peptidyl-lysine deacetylation(GO:0034983) regulation of skeletal muscle fiber development(GO:0048742)
0.0 0.3 GO:1900262 regulation of DNA-directed DNA polymerase activity(GO:1900262) positive regulation of DNA-directed DNA polymerase activity(GO:1900264)
0.0 0.7 GO:0051988 regulation of attachment of spindle microtubules to kinetochore(GO:0051988)
0.0 0.5 GO:0043252 sodium-independent organic anion transport(GO:0043252)
0.0 0.7 GO:0016226 iron-sulfur cluster assembly(GO:0016226) metallo-sulfur cluster assembly(GO:0031163)
0.0 0.2 GO:0045654 positive regulation of megakaryocyte differentiation(GO:0045654)
0.0 0.1 GO:1903026 negative regulation of CREB transcription factor activity(GO:0032792) negative regulation of RNA polymerase II regulatory region sequence-specific DNA binding(GO:1903026)
0.0 0.1 GO:0031117 positive regulation of microtubule depolymerization(GO:0031117)
0.0 0.2 GO:0045719 negative regulation of glycogen biosynthetic process(GO:0045719) negative regulation of glycogen metabolic process(GO:0070874)
0.0 0.3 GO:0021891 olfactory bulb interneuron development(GO:0021891)
0.0 0.5 GO:0034453 microtubule anchoring(GO:0034453)
0.0 0.3 GO:0007076 mitotic chromosome condensation(GO:0007076)
0.0 0.2 GO:0031119 tRNA pseudouridine synthesis(GO:0031119)
0.0 0.8 GO:0031114 negative regulation of microtubule depolymerization(GO:0007026) regulation of microtubule depolymerization(GO:0031114)
0.0 0.1 GO:0046598 positive regulation of viral entry into host cell(GO:0046598)
0.0 0.1 GO:0015808 L-alanine transport(GO:0015808)
0.0 0.1 GO:0048133 germ-line stem cell division(GO:0042078) male germ-line stem cell asymmetric division(GO:0048133) regulation of branching involved in mammary gland duct morphogenesis(GO:0060762) germline stem cell asymmetric division(GO:0098728)
0.0 0.1 GO:1904565 response to 1-oleoyl-sn-glycerol 3-phosphate(GO:1904565) cellular response to 1-oleoyl-sn-glycerol 3-phosphate(GO:1904566)
0.0 0.2 GO:0022027 interkinetic nuclear migration(GO:0022027) astral microtubule organization(GO:0030953)
0.0 0.1 GO:1902268 negative regulation of polyamine transmembrane transport(GO:1902268)
0.0 0.2 GO:0030150 protein import into mitochondrial matrix(GO:0030150)
0.0 0.3 GO:0035235 ionotropic glutamate receptor signaling pathway(GO:0035235)
0.0 0.2 GO:0042761 very long-chain fatty acid biosynthetic process(GO:0042761)
0.0 0.2 GO:0032287 peripheral nervous system myelin maintenance(GO:0032287)
0.0 0.1 GO:0030174 regulation of DNA-dependent DNA replication initiation(GO:0030174) mitotic DNA replication checkpoint(GO:0033314)
0.0 0.1 GO:0061179 negative regulation of insulin secretion involved in cellular response to glucose stimulus(GO:0061179)
0.0 0.4 GO:0002474 antigen processing and presentation of peptide antigen via MHC class I(GO:0002474)

Gene overrepresentation in cellular_component category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 0.6 GO:0042105 alpha-beta T cell receptor complex(GO:0042105)
0.1 0.7 GO:0033093 Weibel-Palade body(GO:0033093)
0.1 0.6 GO:0097149 centralspindlin complex(GO:0097149)
0.1 0.5 GO:0035749 myelin sheath adaxonal region(GO:0035749)
0.0 0.3 GO:0000796 condensin complex(GO:0000796)
0.0 0.3 GO:0031390 Ctf18 RFC-like complex(GO:0031390)
0.0 0.1 GO:0036488 CHOP-C/EBP complex(GO:0036488)
0.0 0.4 GO:0042612 MHC class I protein complex(GO:0042612)
0.0 0.1 GO:0016222 procollagen-proline 4-dioxygenase complex(GO:0016222)
0.0 0.5 GO:0032279 asymmetric synapse(GO:0032279)
0.0 0.2 GO:0005742 mitochondrial outer membrane translocase complex(GO:0005742)
0.0 0.1 GO:0000235 astral microtubule(GO:0000235)

Gene overrepresentation in molecular_function category:

Log-likelihood per target  Total log-likelihood Term Description
0.1 0.3 GO:0008502 melatonin receptor activity(GO:0008502)
0.1 0.3 GO:0016230 sphingomyelin phosphodiesterase activator activity(GO:0016230)
0.1 0.5 GO:0015277 kainate selective glutamate receptor activity(GO:0015277)
0.1 0.5 GO:0015265 urea channel activity(GO:0015265)
0.1 0.4 GO:0038085 vascular endothelial growth factor binding(GO:0038085)
0.1 0.7 GO:0019865 immunoglobulin binding(GO:0019865)
0.1 0.3 GO:0016176 superoxide-generating NADPH oxidase activator activity(GO:0016176)
0.0 0.5 GO:0015347 sodium-independent organic anion transmembrane transporter activity(GO:0015347)
0.0 0.2 GO:0050786 RAGE receptor binding(GO:0050786)
0.0 0.4 GO:0030881 beta-2-microglobulin binding(GO:0030881)
0.0 0.3 GO:0003689 DNA clamp loader activity(GO:0003689) protein-DNA loading ATPase activity(GO:0033170)
0.0 0.5 GO:0070182 DNA polymerase binding(GO:0070182)
0.0 0.2 GO:0015288 porin activity(GO:0015288)
0.0 0.6 GO:0031078 histone deacetylase activity (H3-K14 specific)(GO:0031078) NAD-dependent histone deacetylase activity (H3-K14 specific)(GO:0032041)
0.0 0.1 GO:0004656 procollagen-proline 4-dioxygenase activity(GO:0004656)
0.0 0.3 GO:0005537 mannose binding(GO:0005537)
0.0 0.1 GO:0015180 L-alanine transmembrane transporter activity(GO:0015180) L-proline transmembrane transporter activity(GO:0015193) alanine transmembrane transporter activity(GO:0022858)
0.0 0.5 GO:0017134 fibroblast growth factor binding(GO:0017134)
0.0 0.1 GO:0008073 ornithine decarboxylase inhibitor activity(GO:0008073)
0.0 0.1 GO:0035727 lysophosphatidic acid binding(GO:0035727) lysophosphatidic acid receptor activity(GO:0070915)
0.0 0.1 GO:0008140 cAMP response element binding protein binding(GO:0008140)
0.0 0.2 GO:0034236 protein kinase A catalytic subunit binding(GO:0034236)
0.0 0.2 GO:0009982 pseudouridine synthase activity(GO:0009982)